Perona

Perona (ぺロナ) is girl dressed in a 'gothic lolita' fashion that is able to produce ghosts from her body. She is the Zombie Animal and Zombie Ambush Commander of Thriller Bark. She is one of the main major antagonists of the current arc.

Personality
Perona is confident and firm with herself. She dislikes people telling her what to do and dislikes anyone disobeying her will. She was even willing to put Absalom in his place and remind him of where her authourity lies on Thriller Bark.

She apparently likes 'cute' creatures and will only allow someone to work for her if they are cute. Her dream is to create a country where all the 'cute' people are turned into zombie animals that obey only her. She dislikes Kumacy because, although he is cute in appearance, his voice is not, and thus she prefers him to just be quiet.

She seems to be terrified of cockroaches, even to the point of screaming in fear and believing that plastic cockroaches are real ones. Also, she seems somewhat gullible, not doubting for a second that Usopp can lift ten tons.

Abilities
With the use of her Devil Fruit, Perona is able to conjure up ghosts and form a "ghost network" using them. She either conjures up one ghost or splits herself into many ghosts, able to reform whenever she pleases. Once formed, the ghosts are able to split up into even more ghosts. When a ghost passes through a person they are drained of all their will to live leaving them slumped on the ground regretting even being born, rendering them an easy target.

Despite her powers she so far seems to lack any alternative approaches to fight with. When her Horo Horo no Mi powers failed to stop Usopp she herself became negative and continued to persist with using her powers against him even though she had already failed. However, an additional ability that Perona has is the ability to create exploding ghosts that can make holes in stone walls, and a giant ghost that is even more powerful (although its destructive capabilities are not shown). As well, she can create an astral projection of herself, which can't be hurt through any means. She can pass through solid objects in this form, although she can't damage them, and she can change in size. However, while she is doing this, her true body remains unconscious and vulnerable. To negate this, she usually hides before resorting to astral projection.

History
5 years ago, Perona was sleeping while Brook was causing trouble in Thriller Bark. This explains why she never heard of him.

Perona uses her negative ghosts to scout the ship and the nearby area, presumably looking for people whose shadows can be used for Moria and Hogback's experiments. She recently got into a fight with Usopp, who was immune to her negativity ghost attack. During this fight, her faithful servant Kumacy was effectively killed, since his shadow was removed from him, reverting him to a lifeless corpse. Usopp pursued her and figured out the way her ability (creating a copy of herself using astral projection) worked and succeeded in defeating her using a super-sticky glue to keep her in place, plastic cockroaches (which she took to be real ones), the Usopp Golden Pound, an attack involving his new 10-Ton Hammer (actually a balloon). Oddly enough, despite being the leader of a zombie army, Perona is scared fairly easily. She fears Usopp simply because her negative ghosts don't work on him, and she is so terrified that she faints when encountered with cockroaches and Usopp's "10-Ton Hammer" balloon.