Ryu Ryu no Mi, Model: Pteranodon

The Ryu Ryu no Mi, Model: Pteranodon is an Ancient Zoan-type Devil Fruit that allows the user to transform into a pteranodon hybrid and a full pteranodon at will. It was eaten by King.

Etymology

 * Ryū (竜) is Japanese for "dragon" and contained in the Japanese word for "pterosaur", yokuryū (翼竜).

Strengths and Weaknesses
A user of this fruit's power can turn into a pteranodon: a large and powerful, flight-capable, predatory reptile from prehistory. Once transformed, they become able to fly at tremendous speed, covering large distances in a short time, and gain a significant boost to their overall physical abilities, as expectable of a carnivorous Ancient Zoan. Additionally, the full Beast Form as displayed by King, is notably big, several times larger than an average-sized human (though it is worth noting that King is already much larger than normal people in his normal form).



The combination of great (flying) speed, strength, and size conferred by the transformation permits the user to violently push back and cause notable damage to whatever they fly into headfirst, be it a person or massive structure, such as a large ship. The user's pteranodon beak is long, sturdy, and an effective pointed weapon, allowing King to affix himself to smaller targets and then divebomb into structures in order to tear them apart.

Moreover, the user is given the supposedly natural ability of using their very head like a slingshot of sorts, pulling their beak back into their face by pulling back on their head crest, then releasing their grip to shoot out their head forward a bullet, piercing whatever they aim at with incredible force. According to King, this conforms with how pteranodons used to hunt their prey in ancient times.

Like other models of the Ryu Ryu no Mi, the pteranodon transformation gives the user a tough, dragon-esque hide that significantly increases their durability to a level beyond regular Zoans.

No weaknesses are known of this ability beside those that are standard for Devil Fruits.

Usage


King uses his ability primarily for physical—especially aerial—combat, with his ability to fly being highly advantageous for travel. King has proven able to fly at great speeds, in the anime overtaking Sanji in a high-speed aerial chase while the latter was donning his Raid Suit.

In his full pteranodon form, King can fly into objects with immense speed and force, potentially damaging them greatly in the process. When flying into the Queen Mama Chanter, such midair collision (with a flaming kick in the anime) knocked the enormous ship back with enough force to throw off Big Mom herself. Later, King presented a danger to a Raid Suit-wearing Sanji by diving into him with his strong, pointed beak, challenging said suit's defenses to the point that King claimed he would split his foe in half upon impact (something he ultimately failed at due to the suit proving too durable). In addition to this, King's head too can function as a weapon: pulling back at his head crest allows King to shoot out his head like an extremely fast, oversized bullet, which King has noted is how the pteranodons of the ancient past used to hunt their prey.

Otherwise, as a lunarian, King can combine his transformation with his ability to produce fire; he has been seen attacking with multiple fiery projectiles shot from his wings, almost resembling a living fighter aircraft.

When he is in his Human-Beast Form, King is able to use his pteranodon wing-arms like swords, allowing him to release powerful air blades by swinging them. These slash attacks can deal massive damage to whatever they hit, and he can launch them in rapid succession. His wing blades are also strong enough to clash with and even overpower Meito being wielded by a master swordsman. For defense, King can use his durable wings as a shield; doing so, he was able to withstand a powerful whirlwind of slashes without receiving any scratches.

Techniques
King's techniques have names that end with -don, likely to play on "pteranodon".


 * Tankyudon: A melee attack where King flies toward his opponent in his Human-Beast Form, then hits them with a sweeping strike of one of his wings. When used against Roronoa Zoro, the attack overpowered the latter's Ul-Tora Gari technique before blowing the swordsman back with tremendous force, making him bust right through the Skull Dome's outer wall. In the Viz translation, the technique is called Imperial Archer.


 * Tenpuraudon: A powerful melee attack used by King in his full pteranodon form. He pulls back on his head crest with one arm, stretching his face flat and causing his beak to retract into his face. He then lets go of the crest to let his head fling forward at extreme speed, almost like a slingshot and similar to Luffy's Gomu Gomu No Bell technique, over a distance several times the length of his body, in order to strike his opponent with massive force (presumably using his beak to pierce them). When using this technique against Zoro, the swordsman was hardly able to dodge the attack and unable to block, comparing it to a "beam". The name of the attack is a pun on "tempura udon". In the Viz translation, the technique is called Imperial Deep Pride Stake, as a pun on "deep-fried steak" (tempura being itself a deep-fried dish).

Trivia

 * King possessing this fruit's ability and his "Conflagration" epithet evokes a resemblance to the kaiju Fire Rodan, a pteranodon residing in a volcano, which is among the major reoccurring kaiju in the Godzilla franchise.
 * This is the second Ancient Zoan fruit revealed that is not based on a dinosaur, after the Zou Zou no Mi, Model: Mammoth, although unlike the latter, it still falls under the Ryu Ryu no Mi series.
 * Despite not being dinosaurs, pterosaurs are still close cousins to them. The two families are part of the "Avemetatarsalia" clade from which only three families are known to have descended.

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