Musshuru

Musshuru is the main antagonist in the 9th movie. He is Wapol's older brother in the movie and was designed by Oda for it.

Appearance
In conjunction with his powers, the shape of his head and hair resembles a mushroom, with what appears to be a face mask on his eyes, and black beard at his chin. He also wore a regal armor with a violet cape that covers his right side, orange pants, fluffy shoes, and a weasel-skin scarf on his left shoulder.

Overall, Musshuru has a resemblance with his younger brother and late father, but unlike the former, he is tall and thin, and unlike the latter, he is more peculiar in overall appearance.

Personality
Like his brother, he has an odd speech quirk. Where Wapol calls people "kaba" instead of "baka", he calls everyone "Kabayarou", which means "Hippo-head". He also ends a lot of his sentences with "oi." Like many characters in One Piece, he has a unique laughter style: "Mushushushu", a pun on his own name.

Abilities and Powers
Unlike his younger brother, Musshuru is a much more physical fighter. He possesses both incredible agility and fighting capability. He is capable of moving at speeds that could rival CP9's Soru technique. He is able to execute physical blows such as devastating punches that can smash through a castle spire with great ease. Combined with his Devil Fruit powers, Musshuru is quite a formidable opponent to deal with.

Devil Fruit
Musshuru ate the Noko Noko no Mi, a Paramecia-class Devil Fruit that gives him the power to release and control poisonous spores from his body. This ability of his is so dangerous as the spores by the themselves emitted normally from his body are fatal, and enough to destroy the population of Drum Island. He has crafted several peculiar ways of manipulating these spores. The spores can also be used non-poisonously, as several of his attacks did not poison his opponents on impact.

Though the poison is deadly, it has an antidote to it, and can easily be neutralized by fire. This also makes Musshuru himself very weak against fire, as seen when he was set ablaze, he shrieked in extreme pain.

History
After an incident when he released his spores several years ago, he was exiled from Drum to the Land of Fire, where his spores were nullified by the flame. As Blackbeard invaded Drum, Wapol left to recover his brother hoping his power could assist him in defeating Blackbeard. However, after he left he wishes to use Musshuru's spores to kill off all the people of Drum replacing them with his soldiers.

Thus he accompanies Wapol back to Drum Island only to run into unexpected resistance in the form of Luffy and Chopper at Drum Castle. He fights Luffy, at first evenly match against him, then managing to gain the upper hand by paralyzing him with his spores. Chopper doesn't fare much better but refuses to give up. Before Musshuru can give the final blow he stopped by the arrival of the rest of the Straw Hats who cure Luffy with an antidote. Bringing him back into the fight as Luffy activates Gear 2, Luffy becomes more than a match for Musshuru, finally beating him by using Jet Bazooka and crashing him into the castle's highest tower where Wapol happens to be with his Blitzkrieg Cannon.

Wapol then eats Musshuru along with the cannon, and fuses into a hybrid form with the intention of firing a poison bomb to destroy all of the island. However Chopper manages to knock him into the roof and, despite managing to fire the bomb, it's destroyed by the Straw Hats before it can explode. Luffy then finishes him with a Gear 3 Gigant Pistol that sends them flying from the island.

Trivia

 * His name sounds like the Japanese pronunciation of "Mushroom", matching his hair and spore-based ability.