Fish-Men

Fishmen are one of two species that dominate the seas (the other being the Merfolk). In terms of leadership, none are mentioned, however Fisher Tiger, Jinbei and Arlong were said to have the highest respect amongst their people.

Appearance
Appearance-wise, they are more "fish-like" than merfolk, looking like a cross between a man and a fish of the ocean such as an octopus, manta ray, or sawfish; however, they still have legs.

Biology
They have their gills near their necks which only work when they are in water. On land they can breathe with their lungs like humans do. However, their gills cannot breathe in air, like a normal fish cannot breathe in air through their gills.

Even though Fishmen are part fish, they are still mammals and give birth to live young.

As was revealed during the Davy Back Fight arc with Big Pan Fishmen can also breed with Giants, producing a hybrid who has biological abilities of a Fishmen and a huge body of a Giant (but still smaller than a full one, as related by Robin), they are known as Wotan. Although never shown it's clear that Fishmen and humans can also breed the same way Merfolk and humans can breed.

Inter-species Relationships
Despite their beliefs, usually they get along well with the Merfolk who share the opposite view on life, however it is not unknown for Fishmen to take advantage of the Merfolk's nature.

A stereotypical Fishman considers humans an inferior, weaker species. However, this does not apply to all Fishmen: for example, the pufferfish shipwright Tom cared and looked after two human children (Iceburg and Franky) and even protected them from the actions of Spandam and CP5, and even the Shichibukai, Jinbei, considers the Whitebeard Pirate, Portgas D. Ace an equal and holds the Whitebeard Pirates as a group in high regard because of Whitebeard protecting Fishman Island; he would later place Ace's younger brother, Monkey D. Luffy, under the same treatment. Fisher Tiger was noted to hate humans, but not discriminate when it came to slaves, thus when he broke into the holy land to free his fellow fishmen had also freed slaves of other races in the process. Boa Hancock was noted to have called Fisher Tiger "that man", which shows the respect and gratitude towards the debt she owed him for freeing her.

On the opposite end of the spectrum, human relationship to the Fishmen was defined with their definition of both them and merpeople as "fish" instead of "men". This lead to centuries of discrimination as the Fishmen fought to prove their worth to the world. Only two hundred years ago did the World Government begin to bridge the gap, making an alliance with Fishman Island to remove the problems between the two sides. While attempting to fix the problem, centuries of bad relations remained: many humans continued to see Fishmen as freaks or, in the case around Sabaody Archipelago, slaves for the Tenryuubito. Such bad relations between the races influence certain sects of fishmen, these fishmen would even go so far as to return to the humans what had been done to them. This was seen once in the series with Arlong Park in East Blue where Arlong attempted to enslave the humans as they had enslaved his men and he succeeded in doing so for 8 years.

Overall Strength
Do you know the difference between you and I? Its our species! This is the strength of the Fishmen species from birth. Heaven gave us the power to separate us from you. So we are different! Humans are lower and weaker since the day they were born.

They are ten times as strong as normal humans from birth and this difference is magnified underwater, where a human's power is cut in half (while Fishmen are completely unhindered, and perhaps even strengthened). Jinbei himself noted that on dry land he had little power to offer, which considering the strength that even a fishman as powerful as he has speaks volumes about the potential of any fishman in the water compared to being on dry land. As a whole, in the midst of battle, the Fishman race can be considered unstoppable in their natural habitat - the sea. As well as being powerful they can still talk under water and cannot drown. However breathing air into their gills renders them suffering like a normal fish would, as while they can use their gills to breathe in water, they have to switch back to their lungs to breathe in air.

Their homeland of Fishman Island is also where they learn Fishman Karate, a number of fishmen have displayed skill in this fighting style already within the storyline. This fighting style in particular has both land and sea based attacks, a style suited to the fishman's advantage in the water, but does not leave them completely unable to defend themselves on land when they have to. Their most effective weapon is the sea itself as the most powerful can take a few drops of water and hurl it so hard the impact is as hard as a bullet. To this extent, at the hands of a powerful fishman, even a small splash of water becomes a lethal weapon. As expected they are excellent swimmers they are capable of swimming through the water at a great speed and force, their speed however is no match for the Mermen who are the fastest.

Some species of fishmen have different abilities. Hatchan's six arms gave him a huge advantage over most fighters with just two arms as well as this he can spit ink. Chew as an archerfish could shoot water from his mouth like a bullet. Arlong as a sawfish Fishman could regrow his teeth after they broke. Though only a half breed, Big Pan inherited slippery skin from his mudfish fishman heritage.

Like Giants they have yet to display the use of skills such as Haki and the need for a Devil Fruit ability seems pointless for their race leaving them mostly a physically reliant race like the giants. However, their water based abilities prove most helpful considering that in water Devil Fruit users are rendered powerless completely, the Devil Fruit Kage Kage no Mi in particular has little advantage at all against a Fishman who knows its fatal weakness. Arlong commented on Nami's navigation skills, stating that they can learn about navigation very easily, but they cannot find a good navigator. This is remains unexplained as to why they are particular weak in this area but does indicate their race has an overall weakness in skillful navigation. Arlong himself in particular came to admire Nami's own skills, and in a rare act considered her skills on par with the Fishmen "superiority".

Because of their strength, they appear in One Piece on a regular basis. However, Jinbei was noted to have acknowledged the strength of Sea Kings on one occasion, thus proving even their race is aware of the dangers Sea Kings serve in the grand line.

Beliefs
Their view of life under the sea is dominated by their views that they should rule over all other fish in the sea.

Translation and Dub Issues
The kanji for Fishmen (魚人) is the reverse of the name for Mermen (人魚). They are often incorrectly referred to by fans, and the English dub made by 4Kids, as Mermen for that reason. As a result Mermaids were thought by some to be the female of the species while fishmen were the male, although this is not the case.

Trivia

 * Originally Oda planned on introducing Fishmen as far back as Chapter 3. However, due to his editor he then put it back, ending up introducing the concept during the Arlong arc instead.
 * So far, there isn't a single known fishman who joined the Marines.
 * Octopako is the only fishwoman to be seen in the series so far.