"Z's Ambition Arc - The Little Giant Lily!" is the 575th episode of the One Piece anime.
The Straw Hats find themselves in a middle of an infamous sea in the New World known for its mirages and outrageous weather patterns. After surviving the night, the Straw Hats began to enjoy a relatively calm but hot day at sea only to find that all their food is mysteriously disappearing from right under their noses. Luffy is determined to catch the thief taking his food.
Surrounded by a fleet of Marine battleships, the Straw Hats prepare to fight the overwhelming odds but Nami points out that they were actually mirages. She informs them that they are in the Maubeugemour Sea where dangerous weather phenomena occur and the Straw Hats are in the middle of it. The Straw Hats managed to counter nearly everything the sea throws at them, from humongous rain drops which quickly turned into hail, from snowfall the size of an avalanche and even the very sea splitting into a gorge. After surviving the vicious sea, the Straw Hats enjoy a peaceful but hot day at sea. Sanji prepared some meals for the crew made from whats left of the giant fish Luffy caught. However, Sanji sees that the food he cooked is gone, so he blames it on Luffy, Usopp, and Chopper. Chopper thought the sea must be haunted until he hears a voice coming from the soup pot which was now empty. Suddenly, the food in the kitchen began to be eaten very quickly, leading to a hunt for whatever is eating their food. Sanji, having a bad feeling check the pantry only to find all their food supply gone. After a brief scuffle on deck, Luffy finds the person who ate the food and it turns out to be a miniature woman. Realizing that there is no escape she prepares to fight back and grew to the size of a giant scaring half of the crew. She attempts to strike Luffy but he blocks her weapon and with his superior strength threw her in the sea where she was rescued by Sanji before she drowned.
Thanking them for saving her even though she ate their food, she introduces herself as Lily Enstomach. She turns out to be a giant, but ate a Devil fruit that forced her to be small. She tells them that she was trying to save her father who was being sent to Impel Down for stealing food from the world Government and secretly heard that the ships escorting her father were going to pass through the Maubeugemour Sea. She explained that her father is Panz Fry, a giant cook in the New World famous for his Volcano cooking and giving the stolen food to starving people. Luffy then decided to help her save her father because he wants to try her father's cooking.
On the Neo Marines' ship, Ain calls Shuzo and asks why he was not there with the rest of the fleet. Shuzo tells Ain he is going after Lily's father to kill him so he can replace her as Z's second in command. Back with the Straw Hats, they began to see Marine ships coming their way who have Lily's father captive, so Luffy declares to do battle against the Marines.
- This is the first episode of the Z's Ambition Arc and of the Dressrosa Saga.
- The Neo Marines appear.
- Ain and Binz are shown.
- Shuzo wants to be Z's second in command.
- Lily Enstomach appears on the Thousand Sunny and raids the kitchen. Her ship was destroyed in the unpredictable storms of the Maubeugemour Sea and she took to riding around in a bottle floating in the wreckage, until a whale caused the bottle to land on the Sunny and shatter, destroying her remaining means of transport.
- Lily and her father are giants who steal food from the government and Marines.
- Having eaten the Mini Mini no Mi, Lily is the second known non-canon giant to eat a Devil Fruit. The first one was Blyue who also ate the Mini Mini no Mi.
- The Straw Hat Pirates change their clothes due to getting soaked by the storms in the Maubeugemour Sea and the weather then easing up but turning hot, causing the crew to don summer-appropriate attire. The manga had them change into winter-appropriate outfits when they arrived at a partially frozen island.
- This episode does not contain the New World opening narration.
- A short prologue is seen before the narrator summarizes the previous episode.
- The circumstances the Straw Hat Pirates find themselves in- surrounded by a fleet of Marine warships in stormy waters and luffy using Gigant Pistol against them- parallel the scene in the opening "We Go!" where they are fighting against a fleet of Marines warships under similar weather and end up boarding the ships and using their various abilities and techniques in battle. However, unlike in the opening, the fleet turns out to be mirages caused by freak weather patterns.
- When giant raindrops fall from the sky, as most of the crew is seen kneeling after being knocked over from the rainfall, an animation error occurs- Robin's shoes are briefly missing. It appears as though they fell off, but is evidently not so when the sea parts and she is clinging to the side of the cabin and her shoes are still on.
- Franky dons a rhinoceros beetle hairstyle.
- The preview for the next episode is very brief, only having Luffy state the title of that episode.