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{{Featured Article|# = 161st}}
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{{Featured Article|#=161st}}
 
{{Char Box
 
{{Char Box
| colorscheme = ThrillerBarkColors
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| colorscheme = KuraiganaIslandColors
| image = {{Switch infobox
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| switch = timeskip
|manga pre=[[File:Perona Manga.png|250px|Perona]]
 
|anime pre=[[File:Perona Infobox.png|250px|Perona]]
 
|manga post=[[File:Perona Manga Post Timeskip Infobox.png|250px|Perona]]
 
|anime post=[[File:Perona Anime Post Timeskip Infobox.png|250px|Perona]]}}
 
 
| jname = ペローナ
 
| jname = ペローナ
 
| rname = ''Perōna''
 
| rname = ''Perōna''
| first = [[Chapter 443]]; [[Episode 338]]{{qref|name=Ghost intro|chapter=443|episode=338|text=Perona appears as a ghost before the Straw Hats.}}
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| first = [[Chapter 443]]; [[Episode 338]]{{Qref|name=Ghost Intro|chap=443|page=15|ep=338|Perona appears as a ghost before the Straw Hats.}}
| affiliation = [[Thriller Bark]] (former); [[Mysterious Four]] (former)
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| affiliation = [[Dracule Mihawk]];<br />[[Thriller Bark Pirates]] ([[Thriller Bark Pirates#Organization|Mysterious Four]]){{Qref|name=Enigma|chap=455|page=11-13|ep=349|Perona is identified as one of the Mysterious Four.}}
| occupation = Wild Zombies and Surprise Zombies Commander (former)
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| occupation = [[Pirate]];<br />[[Zombie#Wild Zombies|Wild Zombies]] and [[Zombie#Surprise Zombies|Surprise Zombies]] Commander{{Qref|name=Perona Intro|chap=449|page=18|ep=343|Perona is formally introduced.}}
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| origin = [[West Blue]]{{Qref|name=vivre card|data=vivre card|card=0473|Information about Perona is revealed.}}
| birth = June 7th<ref> [[One Piece Blue Deep: Characters World]] Page 157, Perona's birthday is given</ref>
 
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| residence = [[Kuraigana Island]] (former);<br/>[[Thriller Bark]] (former)
| epithet = {{Nihongo|"Ghost Princess"|ゴーストプリンセス|Gōsuto Purinsesu}}
 
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| epithet = {{Nihongo|"Ghost Princess"|ゴーストプリンセス|Gōsuto Purinsesu}}{{Qref|name=Perona Intro}}
| jva = [[Wikipedia:Kumiko Nishihara|Kumiko Nishihara]]
 
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| age = 23 (debut)<br />25 (after timeskip){{Qref|name=sbs82|sbs=82|page=102|Perona's pre and post timeskip ages and height are revealed.}}
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| birth = June 7th{{Qref|data=blue deep|page=157|Perona's birthday is revealed.}}
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| height = 160 cm (5'3"){{Qref|name=sbs82}}
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| blood type = XF{{Qref|name=vivre card}}
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| jva = [[wikipedia:Kumiko Nishihara|Kumiko Nishihara]]
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| Funi eva = [[wikipedia:Felecia Angelle|Felicia Angelle]]
 
| dfbackcolor = E8E8E8
 
| dfbackcolor = E8E8E8
 
| dftextcolor = DF3982
 
| dftextcolor = DF3982
Line 22: Line 24:
 
| dftype = [[Paramecia]]
 
| dftype = [[Paramecia]]
 
}}
 
}}
'''"Ghost Princess" Perona''' is the former Wild Zombies and Surprise Zombies Commander of [[Thriller Bark]] before the collapse of [[Gekko Moriah|Gekko Moriah's]] [[zombie]] army.{{qref|name=Perona intro|chapter=449|episode=343|text=Perona was finally shown in person.}} She is able to produce ghosts from her body through the powers of the [[Horo Horo no Mi]]. She was one of the major antagonists of the [[Thriller Bark Arc]], being one of the [[Thriller Bark|Mysterious Four]], but has since begrudgingly cooperated with [[Roronoa Zoro]].
 
 
== Appearance ==
 
Perona is drawn in a rather different style from the usual, with big round eyes. Her hair is light pink colored and very long. Her feet are usually seen with her toes pointing inwards in a 'pigeon-toed' manner, whether or not she actually has the [[Wikipedia:pigeon toe|medical condition]] is unknown.
 
   
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'''"Ghost Princess" Perona'''{{Qref|chap=925|page=2|ep=|Perona's name is romanized.}} is the former [[Officer|commander]] of the [[Wild Zombies]] and [[Surprise Zombies]] of [[Thriller Bark]] prior to its collapse.{{Qref|name=Perona Intro}} She was one of the [[Thriller Bark Pirates|Mysterious Four]]{{Qref|name=Enigma}} and one of the major antagonists of the [[Thriller Bark Saga]].
Her looks are inspired by the '[[Wikipedia:Lolita fashion|Gothic Lolita]]' style, with black, white and bright red. Perona's usual hair style has it tied in two high pigtails with somewhat flower-shaped black and white hairpins, and she always wears a red crown with a cross on top. Her tights cover her entire legs and wears a mini-skirt and red buckled boots to match. Her stomach was exposed. She is always wearing red lipstick and a thick layer of eyeliner. She is often seen carrying around a red umbrella shaped somewhat like a cute version of a mini devil before her battle with Usopp.
 
   
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After the defeat of [[Gecko Moria]]'s army and during the timeskip, she begrudgingly helped{{Qref|name=Treating Zoro|chap=524|page=12|ep=421|Perona treats Zoro's injuries after finding him.}} and cooperated with [[Roronoa Zoro]] while in [[Dracule Mihawk]]'s home on [[Kuraigana Island]].{{Qref|name=Preparation|chap=597|page=5|ep=515|Perona prepares to help Zoro heal so he can begin his training.}}
In Hogback's flashback, she is seen as a child. While her crown is visible, the rest of her outfit is hard to see, however she seemed to be wearing a dress that gave her a childish appearance.
 
   
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==Appearance==
After the two year timeskip, Perona has a more mature look than before. Perona is seen on Sabaody Archipelago wearing a long black strapless dress with lavender ruchings, red high heeled boots and a big black top hat with a white floral decor. She has a pink bat tattoo on her left bicep as well as a watch on her left wrist and wears pink nail polish on her fingernails. She still has her signature umbrella and also carries a stuffed animal with her that very much resembles [[Kumashi]]. She also has a different hairstyle now; before the timeskip, it was in two straight pigtails on the side of her head, now her hair is at the back of her head in at least four spiral braids.
 
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Perona is a short young woman with unusually big, round eyes, and she has long, light pink hair that she keeps tied into two pigtails with black and white flower hairpins. She wears a red crown with a black cross on her head, as well as red lipstick and a thick layer of black eyeliner. Her white long-sleeved three different black heart shapes shirt was short enough to expose her belly. She wears a red mini cape with a pink ribbon on front. Her black and white striped tights completely cover her legs, and she wears a mini-skirt and a red and gold belt with a heart belt buckle, as well as red buckled boots. She usually keeps her toes pointing inwards in a pigeon-toed manner and carried around a red parasol shaped like a cute version of a mini devil prior to her battle with [[Usopp]].{{Qref|name=Perona Intro}}
   
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As a child, Perona carried the same parasol and wore the same crown and hairpins but wore her hair in much thicker pigtails and wore a simple, knee-length red dress with a white wave pattern and dots around its center, as well as smaller red boots. She also carried a ghost doll around on a leash and was accompanied by a bear named [[Kumae]].{{Qref|name=Child|sbs=66|movie=12|page=182|Perona is drawn as a child.}}
=== Gallery ===
 
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{{Gallery|height=150|width=150|captionalign=center|position=center
 
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When first meeting [[Hogback]] ten years prior to her encounter with the [[Straw Hat Pirates]], Perona wore the same crown, but the rest of her outfit was not clearly visible, though she seemed to be wearing a similar dress.{{Qref|name=Recruiting Hogback|chap=468|page=9|ep=363|Perona accompanies Moria to Hogback's recruitment.}}
|Kid Perona.png|Perona as a kid.
 
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|Early Moriah.png|Early Perona and her doll (Bottom left corner).
 
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After the two-year timeskip, Perona has a more mature look than before, wearing a long black strapless dress with lavender frills, red high heeled boots and a big black top hat with white floral decor. She has a pink bat tattoo on her left bicep and a watch on her left wrist and wears pink nail polish on her fingernails. She still has her signature parasol, and also carries a stuffed animal resembling [[Kumashi]] with her. Additionally, she now wears her hair in multiple spiral braids on the back of her head.{{Qref|name=Post-Timeskip|chap=602|page=4|ep=522|Perona appears on the Sabaody Archipelago after the timeskip.}}
|Perona Pirate Warriors 2.png|Perona in ''[[One Piece: Pirate Warriors 2]]''
 
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|Perona Berry Match.PNG|Perona in ''[[One Py Berry Match]]''.
 
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While reading about the Straw Hats' actions on [[Dressrosa]] and resulting new bounties, she wore a collared shirt with short, ruffled sleeves and a short dark tie hung loosely around her neck. She wore short frilly skirt with a line of crosses. She had short boots with bows on the front. She also had a large [[straw hat]] with a bow and large crown on top of it.{{Qref|name=chap826|cover=826}}
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===Gallery===
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====Main Series====
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{{Gallery|height=140|width=105|captionalign=center|position=center
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|Perona Pre Timeskip Portrait.png|A close up of Perona's face before the timeskip.
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|Perona Portrait.png|A close up of Perona's face after the timeskip.
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|Perona Post Timeskip Without Hat.png|Perona after the timeskip without her hat on.
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|Perona's Manga Color Scheme.png|Perona's color scheme in the manga before the timeskip.
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|Perona Post Timeskip Color Scheme.png|Perona's color scheme in the manga after the timeskip.
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|Perona Digitally Colored Manga After Timeskip.png|Perona's post-timeskip appearance in the [[Digitally Colored Manga]].
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|Perona as a kid.png|Perona as a kid.
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|Perona 826.png|Perona's outfit during [[From the Decks of the World: The 500,000,000 Man Arc|The 500,000,000 Man Arc]].
 
}}
 
}}
   
== Personality ==
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====Non-Canon====
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{{Gallery|height=140|width=105|captionalign=center|position=center
Perona is confident and firm with herself. She dislikes people telling her what to do and dislikes anyone disobeying her will. She was even willing to put [[Absalom]] in his place and remind him of where her authority lies on Thriller Bark. Despite this, she is normally loyal to [[Gekko Moriah]] and apparently sees him as a way of protection, as she is seen crying to rejoin him. In the manga, she normally speaks using a very masculine language, despite her looks. After Zoro appears, she tends to his wounds, displaying a willingness to befriend even those she sees as a foe. It also shows that she is not completely incapable of being kind to others, despite how cruel she can seem at times.
 
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|Perona Age 13.png|Perona at age 13.
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|Perona Wearing a Coat.png|Perona wearing a coat.
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|Perona Without Her Mini Cape.png|Perona without her mini cape.
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|Perona Boss Luffy Historical Special.png|Perona in "[[Boss Luffy Historical Special]]".
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}}
   
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====Video Games====
She apparently likes cute creatures and will only allow someone to work for her if they are cute. Her dream is to create a country where all the cute people are turned into zombie animals that obey only her. She disliked Kumashi because, although he is cute in appearance, his voice is not, and thus she prefers him to just be quiet. Her desire for cute things gives her a slight childish outlook on some issues. She is very playful, threatening to burst Usopp's heart when she was in her ghost form. Usopp nearly died of the fear itself, and got enraged when Perona simply laughed it off, saying she could do no such thing.
 
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{{Gallery|height=140|width=105|captionalign=center|position=center
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|Perona Pirate Warriors 2.png|Perona in ''[[One Piece: Pirate Warriors 2]]''.
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|Perona DLC Pirate Warriors 2.png|Perona's Okuni DLC outfit in ''One Piece: Pirate Warriors 2''.
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|Perona Wang Yuanji Pirate Warriors 2.png|Perona's Wang Yuanji DLC outfit in ''One Piece: Pirate Warriors 2''.
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|Perona Thriller Bark Warriors 2.png|Perona's Thriller Bark DLC outfit in ''[[One Piece: Pirate Warriors 3]]''.
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|Perona One Py Berry Match.png|Perona in ''[[One Py Berry Match]]''.
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|Perona Super Grand Battle X.png|Perona in ''[[One Piece: Super Grand Battle! X]]''.
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|One Piece Burning Blood Swimsuit Perona (Artwork).png|Artwork of Perona in her ''Swimsuit'' from ''[[One Piece: Burning Blood]]''.
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|Perona Burning Will.png|Perona in ''[[One Piece: Burning Will]]''.
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|Perona Thousand Storm.png|Perona before the timeskip in ''[[One Piece Thousand Storm]]''.
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|Perona timeskip Thousand Storm.png|Perona after the timeskip in ''One Piece Thousand Storm''.
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|Perona Summer Style Thousand Storm.png|Perona (Summer Style) in ''One Piece Thousand Storm''.
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|Perona Ghost Princess Thousand Storm.png|Perona (Ghost Princess) in ''One Piece Thousand Storm''.
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|Perona Wedding Thousand Storm.png|Perona (Wedding) in ''One Piece Thousand Storm''.
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|Perona Valentine Thousand Storm.png|Perona (Valentine) in ''One Piece Thousand Storm''.
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|Perona Bounty Rush.png|Perona in ''[[One Piece Bounty Rush]]''.
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|Perona Jumputi.png|Perona in ''[[Jumputi Heroes]]''.
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|Beautiful Ghost Princess Perona Jumputi.png|"Beautiful Ghost Princess" Perona in ''Jumputi Heroes''.
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}}
   
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====Other====
[[File:Perona Screaming In Fear.png|thumb|left|230px|Perona screaming in fear.]]
 
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{{Gallery|height=140|width=105|captionalign=center|position=center
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|Perona Anime Concept Art.png|Perona's concept art from the anime.
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|Kawaii Tokoro.png|Perona on the cover of the disc ''[[One Piece Nippon Judan! 47 Cruise CD]]''.
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|One Piece novel HEROINES episode PERONA.png|Perona in ''[[One Piece novel HEROINES]]''.
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|Placole Perona.png|Perona's wedding dress from [[Placole Wedding Collaboration Dresses]].
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|Perona Everyday.png|Perona in ''[[One Piece Everyday]]''.
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}}
   
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==Personality==
She seems to have katsaridaphobia, a fear of cockroaches, even to the point of screaming and crying. She is somewhat naïve, not doubting for a second that Usopp can lift ten tons.
 
   
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[[File:Perona's Shocked Face.png|thumb|left|210px|Perona's face fault shocked at Ussop.]]
Like many of the One Piece cast, she has her own [[unique laugh]] that uses the word Horo, which is also part of her Devil Fruit, the [[Horo Horo no Mi]] (i.e Horo-horo-horo-horo-horo!).
 
   
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Perona has a very young, childlike and immature personality; frequently acting selfishly and on her own whims.{{Qref|chap=451|ep=345|page=6|Perona decides to oversee the looting of the Thousand Sunny herself.}}{{Qref|name=Fleeing Thriller Bark|chap=471|ep=366|page=6|Perona decides to flee from Thriller Bark with all the treasure.}} She dislikes people telling her what to do{{Qref|chap=597|ep=515|page=5|Perona angrily tells Mihawk to not order her around.}} and disobeying her will.{{Qref|chap=454|ep=348|page=9|Perona is irritated by Kumashi continuing to speak after she told him not to.}} Despite this, she is normally loyal to [[Gecko Moria]]{{Qref|name=Defeating Straw Hats|chap=460|ep=354|page=11|Perona declares that she will take out the Straw Hats by herself after learning that they have discovered the zombies' weakness to salt.}} and apparently sees him as a father-like role model, as she was crying to rejoin him after being sent to [[Kuraigana Island]].{{Qref|chap=524|ep=421|page=10|Perona expresses her desire to return to Thriller Bark.}} After Zoro appeared on Kuraigana as well, she tended to his wounds, displaying a willingness to befriend even those she formerly knew as a foe and demonstrating that she can be kind to others, despite her normally cruel demeanor.{{Qref|name=Treating Zoro}}
After the timeskip, she has become more mature, although she was still seen carrying a teddy bear around indicating she still held onto many of her past values.{{-}}
 
   
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She is very fond of cute things and will only allow someone to work for her if they are cute.{{Qref|name=Subordinates|chap=449|ep=343|page=12|Perona declares that only cute beings may work for her.}} Her dream is to create a kingdom where all the cute things in the world have been turned into zombie animals that obey only her.{{Qref|chap=456|ep=350|page=10|Perona reveals her dream of creating a kingdom of cute subordinates.}} She dislikes Kumashi because, although he is cute in appearance, his voice is not, and thus she prefers him to just be quiet.{{Qref|name=Shut Up Kumashi|chap=449|ep=343|page=12|Perona is annoyed by Kumashi's deep voice contrasting with his cute appearance.}} Her desire for cute things gives her a slightly childish outlook on some issues.{{Qref|name=Subordinates}} She is somewhat playful during combat, threatening to crush Usopp's heart when she was in her ghost form, which nearly caused Usopp to die from fear, and shortly afterward merely laughing it off when revealing that she could not do so.{{Qref|chap=465|ep=360|page=6-8|Perona tricks Usopp into believing that she will crush his heart.}}
== Relationships ==
 
=== Straw Hat Pirates ===
 
[[File:Perona Scolds Zoro.png|thumb|230px|Perona scolding Zoro.]]
 
   
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[[File:Perona Screaming In Fear.png|thumb|210px|Perona screams in fear at being covered in cockroaches.]]
In the short amount of time she appears, she becomes enemies with one particular member of the [[Straw Hat Pirates]], [[Usopp]]. This is mainly because of him defeating her in battle, handing her the first defeat she had ever suffered. She completely ignored [[Nami]] when she met her later, stating "the long-nose" is her natural enemy, not her. By this, she meant that Nami is an easy opponent for her and she could defeat her only with her Devil Fruit abilities.<ref name="Perona loses">''One Piece Manga and Anime'' - Vol. 49 [[Chapter 473]] and [[Episode 368]], Perona's attitude towards Nami.</ref>
 
   
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Perona has katsaridaphobia, a fear of cockroaches, to the point of screaming and crying when Usopp covered her with fake ones.{{Qref|chap=466|ep=361|page=9|Perona admits her fear of cockroaches upon being covered in fake ones.}} She is also somewhat naive, having watched Usopp seemingly lift a 10-ton hammer without questioning whether or not his feat was genuine, and shortly afterward passing out from fear when he attacked her with it, despite not actually being harmed.{{Qref|name=Defeat|chap=466|ep=361|page=10-12|Perona passes out in fear after being scared by Usopp's tricks.}}
After encountering Zoro on [[Kuraigana Island]], she decided to treat his wounds, and despite the initial quarrels the two develop a relationship of mutual trust by living on the island for two years alongside [[Dracule Mihawk]]. She was also nice enough to bring [[Roronoa Zoro/Personality and Relationships|Zoro]] back to [[Sabaody Archipelago]] and held back the [[Marines]] to allow the Straw Hats to escape.
 
   
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She is very confident in her Devil Fruit powers being able to defeat anyone.{{Qref|chap=473|ep=368|page=15|Perona claims that there is no one whom her powers cannot defeat.}} Even when confronted by the powerful and dreaded [[Seven Warlords of the Sea|Warlord of the Sea]] [[Bartholomew Kuma]], after initially being frozen with fear, Perona quickly regained her confidence and proclaimed that she would defeat him.{{Qref|chap=473|ep=368|page=12-15|Perona is initially frightened by Kuma, but quickly regains her resolve.}} She is also quite stubborn about using them in combat, having continued to use her Negative Hollows against Usopp even after they proved useless due to his negativity because she believed that his immunity was a fluke.{{Qref|chap=462|ep=357|page=2-3|Perona stubbornly continues to use her Negative Hollows against Usopp.}}
=== Thriller Bark ===
 
Perona has followed her master, the [[Shichibukai]] [[Gekko Moriah]], for at least ten years. Her loyalty to him is somewhat contradictory: when she sensed defeat, she attempted to leave and abandon him, but tried to take [[Bartholomew Kuma]] down as a final gift for Moriah; when the [[newspaper]] reported that Moriah died in the [[Whitebeard War]], Perona was deeply saddened.
 
   
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After the timeskip, she has become more mature, although she still carries a teddy bear around, indicating that she is still obsessed with cuteness to some degree.{{Qref|name=Post-Timeskip}} She displayed a grateful side, as she tearfully thanked [[Dracule Mihawk]] for letting her stay on his island.{{Qref|chap=925}}
Perona has another mixed relationship with her teddy bear [[Kumashi|Kumashi (Bearsy)]]. While she thinks of him cute in appearance (which suits her likings), she forbade him from speaking because his voice is not cute. However, she was devastated when Usopp purified Kumashi.
 
   
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Like many others, Perona has her own [[Laughter Style|unique laugh]], which uses the word Horo (e.g., "Horo-horo-horo-horo-horo!"); this is based on the name of her Devil Fruit, the [[Horo Horo no Mi]].{{Qref|chap=449|ep=343|page=12 & 18|Perona laughs while preparing to join the assault on the Straw Hats.}}
== Abilities ==
 
Being formerly one of the Mysterious Four, Perona had a number of Zombies under her command prior to her defection from Thriller Bark. Her ex-crewmate, Doctor Hogback, acknowledges her value under Moria's employment, stating after the fall of Thriller Bark that they would not manage to get very far without her help.
 
   
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==Relationships==
While she seems to recognize that bandages can help a person who is wounded, she seems to lack expertise in medicine. As seen from her treatment of Zoro's wounds, she covered him from head to toe in an overabundance of bandages the first time she found him. However, despite this fault she has common sense or enough knowledge of medical skills, as Zoro recovered from his injuries. She also has at least some knowledge of navigation, seeing how she brought Zoro to Saboady Archipelago all the way from Kuraigana Island.
 
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===Thriller Bark Pirates===
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Perona followed her master, [[Gecko Moria]], for at least ten years.{{Qref|name=Recruiting Hogback}} Her loyalty to him is somewhat complex: when she sensed defeat, she attempted to leave and abandon him{{Qref|name=Fleeing Thriller Bark}} while noting that she had only stayed with him for fun,{{Qref|name=For Fun|chap=472|ep=367|page=18|Perona admits that she had only worked with Moria for fun.}} but tried to take [[Bartholomew Kuma]] down as a final gift for Moria.{{Qref|chap=473|ep=368|page=15|Perona decides to take down Kuma as a parting gift for Moria.}} Despite not clearly showing it, she is said to love Moria like a father since he found her and raised her as a child, and when the [[newspaper]] falsely reported that Moria died in the [[Summit War of Marineford|Whitebeard War]], Perona was deeply saddened.{{Qref|chap=592|ep=509|page=2|Perona grieves over the false report of Moria's death.}}{{Qref|name=SBSChild|sbs=82|page=102|Perona's past with Moria is explained.}} When she learned that Moria is alive two years later, Perona is overjoyed, and she quickly departed Mihawk's castle to go find him.{{Qref|name=c925|chap=925|page=2-3|ep=917|Perona learns Moria is alive and leaves Mihawk.}}
   
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Perona had a mixed relationship with her teddy bear servant, [[Kumashi]]. While she considered him to be cute in appearance, she forbade him from speaking because his voice is not cute.{{Qref|name=Shut Up Kumashi}} However, she was devastated when Usopp purified Kumashi.{{Qref|chap=465|ep=360|page=16|Perona is shocked and angered at the sight of Usopp purifying Kumashi.}}
=== Devil Fruit ===
 
{{see|Horo Horo no Mi}}
 
[[File:Giant Perona.png|thumb|left|200px|Perona's astral projection.]]
 
   
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===Straw Hat Pirates===
With the use of her Devil Fruit, Perona is able to conjure up ghosts and form a "ghost network", using them for surveillance. She either conjures up one or more ghosts or projects her own spirit and splits it into many ghosts able to re-enter her body whenever she pleases (it should be noted that, as a Paramecia user, Perona cannot split her physical body into individual ghosts, only her projected spirit). Using this ability to form her ghost network Perona can see and hear whatever the ghosts can and uses this ability for espionage. Once formed, the ghosts are able to split up into even more ghosts.
 
   
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[[File:Perona Scolds Zoro.png|thumb|left|210px|Perona scolds Zoro.]]
When a ghost passes through a person they are drained of all their will to live leaving them slumped on the ground regretting even being born, rendering them an easy target.<ref name="Ghost Fruit">''One Piece Manga and Anime'' - Vol. 48 [[Chapter 461]] and [[Episode 356]], Perona reveals that she is a Devil Fruit user.</ref> However, when her Horo Horo no Mi powers failed to stop Usopp she herself became negative and continued to persist with using her powers against him even though she had already failed.<ref name="Failing attack">''One Piece Manga and Anime'' - Vol. 48 [[Chapter 462]] and [[Episode 357]], Perona continues to use the same tactics against [[Usopp]], even though they have failed already.</ref>
 
   
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While on [[Thriller Bark]], Perona became enemies with one particular member of the [[Straw Hat Pirates]], [[Usopp]], due to the latter becoming the first person to ever resist the effects of her Negative Hollows{{Qref|chap=462|ep=357|page=2-3|Usopp resists the effects of four Negative Hollows at once, causing the ghosts and Perona herself to become depressed instead.}} and later defeating her in a humiliating manner.{{Qref|name=Defeat}} However, she also cannot help but pity Usopp who lives his life in negativity and attempts to cheer him up, though this may be for her own benefit, as she cannot make him negative unless he was already slightly positive.{{Qref|name=cheerup|chap=461|ep=356|page=11-18|Perona confronts several of the Straw Hats, but is shocked to learn Usopp is immune to her Negative Hollows.}}{{Qref|name=cheerup2|chap=464|ep=359|page=16-19|Perona flees from Usopp before exiting her body in order to begin her counterattack.}} She looked down on [[Nami]] when the latter confronted her later, claiming that she could easily defeat Nami with her Mini Hollows because Usopp was her only natural enemy.{{Qref|chap=473|ep=368|page=11|Perona talks down to Nami.}}
An additional ability that Perona has is the ability to create exploding ghosts that can make holes in stone walls, and a giant ghost that is even more powerful (although its destructive capabilities are not shown). She can also create an astral projection of herself, which cannot be hurt through any means. She can pass through solid objects in this form, although she cannot damage them, and she can change in size. However, while she is doing this, her true body remains unconscious and vulnerable. To negate this, she usually hides before resorting to astral projection.{{-}}
 
   
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After encountering Zoro on [[Kuraigana Island]], she was shocked to see him but decided to treat his wounds as he was the only other person there at the time.{{Qref|name=Treating Zoro}} Despite their initial quarrels,{{Qref|chap=524|ep=421|page=12|Perona and Zoro argue over Perona taking away Zoro's swords.}} the two developed a relationship of mutual trust by living on the island for two years alongside [[Dracule Mihawk]].{{Qref|chap=602|ep=522|page=4|Perona interacts with Zoro after bringing him to Sabaody Archipelago.}} She also willingly brought Zoro back to [[Sabaody Archipelago]]{{Qref|name=Transporting Zoro|chap=602|ep=522|page=4|Perona reveals that she brought Zoro back to Sabaody Archipelago.}} and later held back the [[Marines]] in order to allow the Straw Hats to escape.{{Qref|chap=602|ep=522|page=13|Perona uses her Negative Hollows to demoralize a Marine unit.}}
== History ==
 
=== Perona Before Joining Gekko Moriah ===
 
Perona's past is unknown. According to her own words, she joined Moriah merely for fun; it is unknown what that "fun" even meant.
 
   
=== Thriller Bark Arc ===
+
===Dracule Mihawk===
  +
After being sent to [[Kuraigana Island]] by [[Bartholomew Kuma|Kuma]]'s [[Nikyu Nikyu no Mi|abilities]], Perona decided to take residence in the castle where the at the time [[Seven Warlords of the Sea|Warlord]] Mihawk lived.{{Qref|chap=524|ep=421|page=10|Perona begins inhabiting Mihawk's castle.}} Mihawk seemingly agreed or simply did not mind letting her stay with him, though he did not display any hospitality toward her beyond that.{{Qref|chap=592|ep=509|page=2|Mihawk allows Perona to stay in his castle, but does not display any more hospitality than that toward her.}}
[[File:Perona usopp.png|thumb|210px|Perona vs. Usopp.]]
 
   
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It was shown in [[From the Decks of the World: The 500,000,000 Man Arc]] that even after going back to [[Thriller Bark]] to retrieve [[Kumashi]], she returned to Kuraigana Island and was seen helping him with his work in tilling the land, suggesting a good status between them.{{Qref|chap=826}} When she learned of her old master's survival, she was upset that Mihawk did not tell her and nearly left as he told her to be safe while she tearfully thanked him for letting her stay with him.{{Qref|name=c925}}
Perona has been with [[Gekko Moriah|Moriah]] and [[Absalom]] for over 10 years. When she is seen meeting Hogback, she was still a young girl. She was holding a bear that seems to have been later used as source of inspiration to make a real life replica, [[Kumashi]].<ref name="Perona's past">''One Piece Manga and Anime'' - Vol. 48 [[Chapter 468]] and [[Episode 363]], Perona is seen meeting Hogback along with Gekko Moriah and Absalom.</ref>
 
   
  +
==Abilities and Powers==
Five years ago, she was sleeping while [[Brook]] was causing trouble in [[Thriller Bark]]. This explains why she never heard of him. When Kumashi tried to wake her up, she simply told him to shut up and went back to sleep.
 
  +
Being formerly one of the Mysterious Four, Perona had a number of zombies under her command prior to her defection from Thriller Bark.{{Qref|name=Perona Intro}} Her ex-crewmate, Doctor [[Hogback]], acknowledges her value under Moria's employment.{{Qref|chap=460|ep=355|page=11|Hogback admits that they are lucky to have Perona on their side.}}
   
  +
Perona is physically strong, as she was seen carrying Kumashi's body, which is bigger than she is, while levitating with her Devil Fruit powers.
Perona uses her negative ghosts to scout the ship and the nearby area, presumably looking for people whose shadows can be used for Moriah and Hogback's experiments.
 
   
  +
Perona is able to cook, having cooked several times during the two years while she was living with Mihawk.{{Qref|name=c925}}
She got into a fight with [[Usopp]], who was immune to her negativity ghost attack. During this fight, her faithful servant Kumashi was effectively killed, since his shadow was removed from him, reverting him to a lifeless shell. Usopp pursued her and figured out the way her ability (creating a copy of herself using astral projection) worked and succeeded in defeating her using a super-sticky glue to keep her in place, plastic cockroaches (which she took to be real ones), the Usopp Golden Pound, an attack involving his new 10-Ton Hammer (actually a balloon).
 
   
  +
===Medical Knowledge===
[[File:Kuma peronna.png|thumb|left|200px|Perona meets Kuma.]]
 
  +
While she seems to recognize that bandages can help a person who is wounded, she seems to lack expertise in medicine. During the process of treating Zoro's wounds, she covered him from head to toe in an overabundance of bandages the first time she found him.{{Qref|name=Treating Zoro}} However, despite this fault, she has common sense or enough knowledge of medical skills, as Zoro recovered from his injuries.{{Qref|chap=597|ep=515|page=5|Perona is entrusted with ensuring Zoro's wounds heal.}}
   
  +
===Navigation Knowledge===
Oddly enough, despite being the leader of a zombie army, Perona is scared fairly easily. She fears Usopp simply because her negative ghosts do not work on him, and she is so terrified that she faints when encountered with cockroaches and Usopp's [[Usopp's_Arsenal#Hammers|"10-Ton Hammer"]] balloon.<ref name="Perona loses">''One Piece Manga and Anime'' - Vol. 48 [[Chapter 466]] and [[Episode 361]], Perona loses a fight for the first time, against Usopp.</ref>
 
  +
She is a capable navigator, demonstrated when she brought Zoro to Sabaody Archipelago all the way from Kuraigana Island.{{Qref|name=Transporting Zoro}} She later traveled to Thriller Bark and returned back to Kuraigana Island all by herself. After learning Moria was alive, she traveled to all the way to [[Hachinosu]] from Kuraigana Island, going from [[Paradise]] to the [[New World]].
   
  +
===Devil Fruit===
She is later seen, still passed out, surrounded by her zombies while [[Oars]] rampages through Thriller Bark. When she wakes up later, after having been taken by the zombies to another (probably safer) place, she's still confused and scared, thinking that Usopp is still there, and keeps screaming about the hammer and the cockroaches as if she was still talking to him. She's then informed by the zombies that Usopp's gone already and that the damage done to the building wasn't made by the [[Straw Hat Pirates]] like she first thought, but by Oars.
 
  +
{{See|Horo Horo no Mi}}
   
  +
[[File:Horo Horo no Mi Power.png|thumb|210px|Perona's negative ghosts mocking her enemies.]]
Just after they finish talking, Oars' arm broke the wall close to her, hitting a few zombies. Scared, she hid behind a rock that should have been a part of either the wall or the ceiling before. When she hears Oars saying he's looking for a "woman" (referring to Nami), she freaks out and decides to run away from Thriller Bark using the pirates' ship, as she thinks that this is way too much for her and that Thriller Bark is already done for.<ref name="Perona leaves">''One Piece Manga and Anime'' - Vol. 49 [[Chapter 471]] and [[Episode 366]], Perona decides to leave Thriller Bark believing it's not safe anymore.</ref>
 
   
  +
Perona ate the [[Horo Horo no Mi]], a [[Paramecia]]-type [[Devil Fruit]] that allows her to create and control ghosts. The ghosts have the power to drain the morale, self-esteem, and will to live of anyone whom they pass through, described as hollowing out the victim's spirit.{{Qref|name=Reveal|chap=461|ep=356|page=13|Perona reveals the name of her Devil Fruit and explains its power.}}{{Qref|chap=448|ep=342|page=9-10|Franky, Luffy, and Zoro are made negative by the ghosts passing through them.}} She can also create smaller ghosts that explode with enough force to damage stone walls{{Qref|chap=465|ep=360|page=10-11|Perona demonstrates the power of her Mini Hollows}} and larger ghosts that produce an even more powerful blast.{{Qref|chap=466|ep=361|page=5-6 & *|Perona uses the Toku Hollow, and Usopp absorbs the blast with an Impact Dial and uses it to defeat Hippo Gentleman.}}
As she comes upon the ''Sunny'', she is confronted by [[Bartholomew Kuma]]. When she refuses to tell him about where Moriah is, he asked her where she would like to go, and she replies 'somewhere near an old castle engulfed in malice'. Kuma then seems to make her disappear using his [[Nikyu Nikyu no Mi]] ability. When Hogback and Absalom escaped Thriller Bark later, they left without Perona, stating they could not find her anywhere.<ref name="Perona's disappearance">''One Piece Manga and Anime'' - Vol. 50 [[Chapter 486]] and [[Episode 378]], Perona could not be found by her comrades.</ref>{{-}}
 
   
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Her ghosts can also be used to form a "Ghost Network" for surveillance. This allows her to spy on enemies.{{Qref|chap=453|ep=347|page=13|Tararan identifies Perona's surveillance system as her "ghost network".}}{{Qref|name=Surveillance|chap=443|ep=338|page=16|Brook notes the Thousand Sunny is already under surveillance by Perona's ghosts.}} The ghosts can also duplicate themselves into even more ghosts.{{Qref|chap=448|ep=342|page=8|One of Perona's ghosts splits into three ghosts.}}
=== Straw Hat Separation Adventure: Kuraigana Island ===
 
[[File:Perona Finds Zoro.png|thumb|230px|Perona encounters Zoro.]]
 
   
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In addition to creating ghosts, she can make her own ghost leave her body. Her disembodied ghost is completely intangible; she can pass through solid objects and cannot be hit by enemy attacks, although she cannot damage anything or attack her opponents physically.{{Qref|chap=465|ep=360|page=9|Perona demonstrates that all attacks will go through her astral projection.}}{{Qref|chap=465|ep=360|page=6-8|Perona demonstrates her ability to pass through solid matter, but reveals that she cannot physically interact with it.}} Her ghost also has the power to levitate and can change the size of her spectral body.{{Qref|chap=465|ep=360|page=4-5|Perona changes the size of her astral projection.}} However, while she is doing this, her true body remains unconscious and vulnerable.{{Qref|chap=466|ep=361|page=4|Perona panics over Usopp attacking her real body.}}
Kuma honored Perona's request, having sent her to [[Kuraigana Island]], an island filled with dark and spooky ruins, in addition to the malicious castle. Although she has taken up residence in the castle, Perona is not happy because she has no servants or cute animals. As she laments her situation, wishing she was back on Thriller Bark, [[Roronoa Zoro]] then lands nearby her. She goes to investigate and at first she is excited, but then reacts with terror when she recognizes him as one of the [[Straw Hat Pirates|Straw Hats]]. Since Zoro is the only person there, she decides to save him. She brings him to the castle and treats his wounds, but bandages him in a ridiculous manner due to her insufficient medical skills.
 
   
  +
==History==
When Zoro comes around, he finds out that she has taken his swords away to prevent him causing problems for her.<ref name="Perona's location">''One Piece Manga and Anime'' - Vol. 54 [[Chapter 524]] and [[Episode 421]], Perona's whereabouts revealed.</ref> However, some brand of trust seems to have grown between them, as Zoro is later seen with his swords at his side and is casually walking through the ruins of the island as Perona floats overhead (though that makes it clear that Zoro could not have harmed her in any way at that time).
 
  +
===Past===
  +
When she was a child, Perona owned a pet bear named [[Kumae]].{{Qref|name=Child}} At some point in her childhood, she met the pirate [[Gecko Moria]] who took her in and raised her and she eventually saw him as a father figure.{{Qref|name=SBSChild}}
   
[[File:Mihawk_at_home.png|thumb|left|250px|Perona and Mihawk discussing Moriah's fate.]]
+
[[File:Hogback Meets Moria, Absalom and Perona.png|thumb|left|210px|Perona witnesses the meeting between Moria and Hogback.]]
   
  +
Over 10 years ago, she joined Moria's crew merely for fun.{{Qref|name=For Fun}} She and [[Absalom]] were present when Moria met and recruited the genius doctor, [[Hogback]].{{Qref|name=Recruiting Hogback}}
Later, it is revealed that the castle that Perona was sent to is the home of [[Dracule Mihawk]]. She reads the news stating that Moriah had died in the Whitebeard War and starts to cry. Mihawk then tells Perona to cry elsewhere. Perona then scolds Mihawk for his lack of sympathy. Mihawk also says that the news about Moriah's demise lacks credibility, but Mihawk could not give Perona a definite answer if Moriah is alive or not.
 
   
  +
Five years ago, she was sleeping while [[Brook]] was causing trouble in [[Thriller Bark]], which would later cause her to not know what he looked like when he returned.{{Qref|chap=456|ep=350|page=12|Perona reveals that she does not know what Brook looks like.}} When [[Kumashi]] tried to wake her up, she simply told him to shut up and went back to sleep.{{Qref|chap=458|ep=352|page=11|Perona sleeps through the crisis caused by Brook on Thriller Bark.}} Three and a half years ago, Perona came to Moria with news that [[Bartholomew Kuma]] was elected the new [[Seven Warlords of the Sea|Warlord of the Sea]], Moria brushed it off finding his moniker funny but Perona told him that he must be a strong figure to be chosen as a Warlord.{{Qref|chap=1100|page=16|Perona tells Moria that a new Warlord was elected.}}
Later, she helps an apparently injured Zoro read the article about Luffy going to Marineford again by holding it up for him to read. She was complaining that he was taking a long time and that her hands were getting tired.
 
   
  +
===Thriller Bark Saga===
After accepting Zoro's request for training, Mihawk tells Perona to tend to Zoro's wounds. She tells Mihawk not to order her around. Later, Perona asks Zoro about Luffy's message, and the explanation is given.{{-}}
 
  +
====Thriller Bark Arc====
  +
Perona used her Ghost Network to scout out the [[Thousand Sunny]]{{Qref|name=Ghost Intro}} and later reported [[Straw Hat Pirates|the crew]]'s [[bounties]] to Hogback and Absalom.{{Qref|name=Report|chap=447|ep=341|page=7|Perona reports the bounties of the Straw Hats to Absalom and Hogback via one of her ghosts.}}
   
  +
In the Dead Forest, a group of Negative Hollows encountered some of the Straw Hats and, after evading capture, demoralized [[Monkey D. Luffy]], [[Franky]], and [[Roronoa Zoro]].{{Qref|chap=448|ep=342|page=8-10|Perona's ghosts demoralize some of the Straw Hats in the forest.}} As the Night Attack began, Perona recalled her ghosts and rallied her [[Wild Zombies]] and [[Surprise Zombies]].{{Qref|chap=449|ep=343|page=12 & 18|Perona returns to her bodies and rallies her zombies.}} When she left to oversee the looting of the Thousand Sunny, she ordered Kumashi to meet the [[Risky Brothers (Zombies)|Risky Brothers]], who were carrying [[Nami]], [[Usopp]], and [[Tony Tony Chopper]] in coffins, and take them to the Dance Hall.{{Qref|chap=451|ep=345|page=6|Perona orders Kumashi to assist the Risky Brothers.}}
=== Two Years Later ===
 
[[File:Perona&Kumacy.png|thumb|230px|Perona and Kumashi reunited.]]
 
   
  +
On the Thousand Sunny, Perona was frustrated at the lack of treasure and had the food taken from the ship. [[Hildon]] reported to Perona that Moria was gathering the other members of the Mysterious Four because he had captured Luffy and would be debuting [[Oars|Zombie 900]], much to her surprise. Soon afterward, Perona returned to her room, where she found Absalom speaking with Kumashi about the escape of Nami, Usopp, and Chopper. After scolding Kumashi for trying to speak, Perona informed Absalom that Moria was calling them together and explained why he was doing so.{{Qref|chap=454|ep=348|page=7-9|Perona learns of Moria summoning the other members of the Mysterious Four and informs Absalom of it as well.}} In the Mast Mansion, the Mysterious Four discussed Luffy and the Straw Hats, and Perona reported that Nami, Usopp, and Chopper had escaped before yelling at Kumashi when he tried to correct her. When Luffy broke free from his cage, Perona quickly stopped him with her Negative Hollow before watching on as Moria removed Luffy's shadow.{{Qref|chap=455|ep=349|page=11-19|Perona meets with the rest of the Mysterious Four and incapacitates Luffy.}} After hearing Nami, Usopp, and Chopper talking inside Kumashi and believing that it was Kumashi himself doing so, Perona proclaimed that her dream of creating a kingdom full of cute subordinates was closer to fruition with Luffy's shadow extracted. When two Spider Mice appeared and reported that [[Tararan]]'s shadow had been extracted, Hogback noted that it could only have been the "Humming Swordsman", a name which Perona did not recognize. Soon afterward, Perona accompanied the rest of the Mysterious Four to the enormous freezer containing the corpse of [[Oars]].{{Qref|chap=456|ep=350|page=8, 10-12 & 17-19|Perona learns of Brook's return and enters the freezer containing Oars's corpse.}} Perona watched as Moria used Luffy's shadow to revive Oars's corpse. When Oars awoke, Nami, Usopp, and Chopper screamed and fell from Kumashi's back, prompting Perona to yell at Kumashi for hiding the pirates.{{Qref|chap=457|ep=351|page=7-9|Perona witnesses Oars's awakening and learns of the Straw Hats hiding inside of Kumashi.}} Perona continued to chastise Kumashi for hiding the pirates without letting him respond.{{Qref|chap=458|ep=352|page=4|Perona berates Kumashi for hiding the Straw Hats.}}
After Zoro's two year training, Perona took him back to Sabaody Archipelago (solving the mystery of how he managed to arrive first). As Luffy, Zoro, and Sanji made their way to the [[Thousand Sunny]], Perona aided them by fending off the Marines with her Negative Hollow. She then asks what they were still doing loafing around on the archipelago when they should be getting ready to leave. She also scolds Zoro for his ungrateful attitude towards her, and brushes off Sanji, who is showing his infatuation for her, commenting that the chef has some kind of illness. She then informs the Straw Hats that Marine vessels were coming soon. Later, she assisted in the Straw Hats' escape by holding back the second Marine platoon with her Negative Hollows.
 
   
  +
After reporting Oars's current activities to Moria, Perona learned that the Straw Hats had learned of the zombies' weakness to salt and were approaching them, with the General Zombies preoccupied with attending Absalom's wedding, but dismissed Hogback's worry over the zombies' weakness being known and retired to her room before attacking the Straw Hats with her ghosts, planning to weaken them enough to be taken by [[Zombie#Soldier Zombies|Soldier Zombies]]. However, Perona's attack was interrupted by Oars crashing down through the mansion.{{Qref|chap=460|ep=355|page=8, 11 & 15-17|Perona attempts to weaken the Straw Hats with her Negative Hollows so they cannot purify any zombies but is interrupted by Oars.}} After Franky rebuilt the destroyed bridge, he, Zoro, Sanji, and Usopp charged into Perona's room, where Perona confronted them. After explaining the powers granted by her Devil Fruit, Perona sent a Negative Hollow through each of the Straw Hats, but Usopp was unaffected and purified several of her zombies before declaring that he was always negative, shocking Perona, who tried to cheer him up alongside her zombies before preparing to battle him.{{Qref|name=cheerup}}
Much later, she left Sabaody Archipelago and returned to the wandering Thriller Bark, being happy to find [[Kumashi|Kumashi's]] lifeless body there.{{-}}
 
   
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[[File:Perona vs. Usopp.png|thumb|210px|Perona vs. Usopp.]]
== Major Battles ==
 
* Perona vs. [[Usopp]]
 
* Perona vs. [[Bartholomew Kuma]]
 
* Perona vs. [[Marines]]
 
   
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Believing Usopp's immunity to have been a fluke, Perona sent four Negative Hollows through him, but instead became depressed herself alongside her ghosts, to the shock of her zombies.{{Qref|chap=462|ep=357|page=2-3|Perona becomes negative after attempting to overwhelm Usopp with four Negative Hollows at once.}} Soon afterward, Perona fled from Usopp while noting that she could defeat the other Straw Hats if it were not for him. After Kumashi distracted Usopp by attacking him, Perona projected her spirit from her body after hiding and tricked Usopp into believing she was flying while proclaiming that she had come up with a plan to defeat him.{{Qref|name=cheerup2}} Declaring that she had simply been caught up in the moment before, Perona attempted to cheer Usopp up once more before appearing behind him in a giant form. After dodging Usopp's Ageha Ryusei, Perona reverted to her normal size before placing her hands inside of Usopp's chest and pretending that she was going to crush his heart, only to reveal that she could not do such a thing. Usopp attacked her with Rokuren Mamushi Boshi, but Perona demonstrated her intangibility by letting the attack go through her and moving through the floor. When Usopp pointed out that she could not touch him either and demanded to know how she could attack him, Perona summoned some Mini Hollows and demonstrated their explosive power on a nearby wall before attacking Usopp with one as well. After tricking Usopp into running into a pillar, Perona covered his body with Mini Hollows and detonated all of them before gleefully watching Kumashi brutalize Usopp. However, Perona was shocked to see Usopp, who had donned his Sogeking mask, purify Kumashi with a salt ball before fleeing once more. Wounding Usopp with a barrage of Mini Hollow blasts once more, Perona decided to hand him over to Moria, but was surprised when Usopp figured out her trick and used Atlas Suisei to expose the room containing her true body.{{Qref|chap=465|ep=360|page=2-19|Perona battles Usopp and initially overwhelms him, but has her true body exposed.}}
== Anime and Manga Differences ==
 
=== Thriller Bark Arc ===
 
In the manga, when Usopp knocked her out, foam was coming out of her mouth. It was edited out in the anime.
 
   
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[[File:Usopp Defeats Perona.png|thumb|left|210px|Usopp defeats Perona.]]
=== Straw Hat Separation Serial ===
 
[[File:Zoro as Kumacy.png|thumb|left|230px|Perona dresses Zoro up as Kumashi.]]
 
   
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After Usopp deduced how she left her body by becoming a ghost in order to fight him, Perona revealed that she was a fully conscious Hollow instead of merely being a ghost controlled by her body, prompting Usopp to seemingly attack her body in an attempt to destroy it. After the attack apparently missed, a panicked Perona engulfed his body with a Toku Hollow before returning to her own body. Perona detonated the Toku Hollow but was surprised to see the explosion had disappeared as Usopp revealed he had absorbed it with an [[Dials#Impact Dial|Impact Dial]]. After learning that Usopp had actually shot a Torimochi Boshi at her bed in order to trap her body in one place, Perona witnessed Usopp defeat [[Hippo Gentleman]] with a blast from his Impact Dial before covering Perona herself in fake cockroaches with Kurobikari Boshi. Crying and screaming hysterically at the sight of cockroaches, a shocked Perona saw Usopp seemingly lifting a 10-ton hammer and begged him to spare her before passing out in fright as the fake hammer burst apart on impact with her. Usopp celebrated his victory over her as Perona collapsed while foaming at the mouth.{{Qref|chap=466|ep=361|page=2-12|Usopp outmaneuvers and defeats Perona.}}
Also, in the anime version of Zoro's cover story, she displays more of a lonely childlike personality, mostly due to living by herself with no servants. She even tries to force Zoro to be her servant by using her Negative Ghosts to make him swear to be her servant and dresses him up as [[Kumashi]].
 
   
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Later, Perona's zombies found Perona still unconscious but mostly unharmed as Oars rampaged through Thriller Bark.{{Qref|chap=469|ep=364|page=17|An unconscious Perona is found by her zombies.}} Soon afterward, Perona regained consciousness and had to be calmed down by her zombies when she acted like Usopp was still attacking her. Upon seeing the damage done to the mansion by Oars, Perona wondered if the Straw Hats had done this and was shocked to learn that Oars had completely wiped out the General Zombies as well, which caused her to question if they could control him at all. However, the zombies' explanation of what was happening was interrupted by Oars punching them through one of the nearby walls while searching for Nami; witnessing this, Perona was convinced that Oars and the Straw Hats were going to completely destroy Thriller Bark and ordered her remaining subordinates to gather all the treasure and some supplies so they could leave on the Thousand Sunny.{{Qref|chap=471|ep=366|page=4-6|Perona decides to leave Thriller Bark after witnessing the destruction caused by Oars.}}
In the anime, after Luffy's message is revealed, Perona went outside the castle with Mihawk and Zoro. She is seen scolding Zoro to skip his training until his wounds are healed.
 
   
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[[File:Perona Encounters Kuma.png|thumb|210px|Perona meets Kuma.]]
In the anime, Perona's scene with Sanji is extended as the latter is seen beaten up and inflicted with Negative Hollow for freaking out the former. Also, as she uses her ghosts to fend off the marines, she is seen looking out to sea and states that she will find another boytoy, implying that Zoro was one, seemingly disappointed at his departure.{{-}}
 
   
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Observing the battle between the Straw Hats and Oars from the Thousand Sunny, Perona admitted that she only joined Moria for fun and would not stay with him if her life were in danger before telling her subordinates that they would be leaving without delay.{{Qref|chap=472|ep=367|page=18-19|Perona notes why she is leaving Thriller Bark while watching the battle against Oars.}} When Nami confronted her over her taking the treasure and the Thousand Sunny, Perona declared that she was planning to leave Thriller Bark on the Thousand Sunny and prepared to battle while proclaiming that her Mini Hollows should be enough to defeat Nami. However, she was interrupted by the appearance of [[Bartholomew Kuma]], whom Perona was terrified at the sight of due to knowing his reputation as a cruel pirate and a Warlord of the Sea. When Kuma asked her where she would like to go on a trip, Perona described a dark, damp island with an old castle filled with malice before realizing that she had gotten caught up in his pace. Resolving to take down Kuma as a parting gift for Moria due to his shadow likely being strong, Perona prepared to attack him with a combination of Negative Hollows and a Toku Hollow, but Kuma suddenly [[Nikyu Nikyu no Mi|made her disappear]] before her attack could connect, to the confusion of those watching.{{Qref|chap=473|ep=368|page=10-16|Perona confronts Kuma, but is effortlessly defeated.}}
=== Return to Sabaody Arc ===
 
When Perona is scolding Zoro at Sabaody, her hat bended in contact with the swordsman. This suggests that she was in her physical form rather than her ghost form, however this doesn't happen in the manga. Similar inconsistencies were shown during the filler scenes at Kuraigana: although it was clearly shown she was in her ghost form, she had some interactions with physical objects, indicating they were simply errors.{{Qref|name=hat|chapter=602|episode=522|text=Perona scolds Zoro.}}{{Qref|name=errors|chapter=|episode=456|text=Although Perona is her ghost form, she fells on the ground raising dust}}
 
   
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When Hogback and Absalom escaped Thriller Bark later, they left without Perona while noting they could not find her anywhere.{{Qref|chap=486|ep=378|page=2|Hogback and Absalom could not find Perona.}}
== Early One Piece ==
 
An early concept of Perona from [[One Piece Green: Secret Pieces]] shows how she was supposed to be a silent, princess-like girl. {{-}}
 
   
== Merchandise ==
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===Summit War Saga===
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====Amazon Lily Arc====
She has been confirmed to feature in the series of Portrait of Pirates figurines, [http://mangafigures.wordpress.com/2011/01/23/ghost-princess-perona-excellent-model-one-piece-neo-dx-portraits-of-pirates-18-scale-pre-painted-pvc-figure/ in her pre-timeskip attire].
 
   
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[[File:Perona Finds Zoro.png|thumb|210px|left|Perona encounters Zoro.]]
== Trivia ==
 
[[File:Nami 4th Thriller Bark Outfit.png|thumb|155px|Perona's personal Jolly Roger on Nami's blouse.]]
 
   
  +
Kuma honored Perona's request, having sent her to [[Kuraigana Island]], an island filled with dark and spooky ruins, in addition to the malicious castle. Although she had taken up residence in the castle, Perona was not happy because she had no servants or cute animals. As she lamented her situation while wishing she was back on Thriller Bark, Perona noticed a disturbance in the forest and discovered that Zoro had been sent to the island as well. Initially terrified by the sight of one of the Straw Hats, Perona eventually decided to save him since he was the only other person there at the time and brought him to the castle, where she treated his wounds by heavily bandaging him. When Zoro regained consciousness, he found out that she had taken his swords away to prevent him from attacking her.{{Qref|chap=524|ep=421|page=10-12|Perona laments her situation and discovers Zoro has been sent to Kuraigana Island as well.}}
*Perona has a personal [[Jolly Roger]], which consists of a skull with swords a cross on either side of the grinning mouth, yellow hair on the sides with a flaming pattern, and the crown that Perona herself wears. [[Nami]] was seen wearing a top with Perona's own personal [[Jolly Roger]] on it when she rejoins the other Straw Hats, most likely having stolen them from Perona's room after disposing of the wedding dress Absalom forced her into.<ref name="Perona's symbol">''One Piece Manga and Anime'' - Vol. 49 [[Chapter 475]] and [[Episode 370]], Nami is seen wearing a top with Perona's skull and crossbones on it.</ref>
 
* In the 4th Japanese [[Popularity Polls|Fan Poll]], Perona is ranked as the 33rd most popular character in One Piece.{{-}}
 
   
  +
====Straw Hat's Separation Serial====
== See also ==
 
  +
Eventually, Perona returned Zoro's swords and followed him through the ruins of the island, indicating that they had developed some level of trust by then.{{Qref|cover=559|ep=456|Perona follows Zoro through the ruins of Kuraigana Island.}}
*[[Dracule Mihawk]]
 
*[[Gekko Moriah]]
 
*[[Kumashi]]
 
*[[Kuraigana Island]]
 
*[[Thriller Bark]]
 
*[[Usopp]]
 
   
== References ==
+
====Post War Arc====
  +
{{Scroll box
 
  +
[[File:Mihawk at Home.png|thumb|210px|Perona and Mihawk discuss Moria's fate.]]
|content=<font size="1.5">
 
  +
<references/>
 
  +
After the [[Summit War of Marineford]], the owner of the castle that Perona was inhabiting, [[Dracule Mihawk]], returned to Kuraigana Island, and Perona began sobbing at the news of Moria's death in the newspaper. When Mihawk told her to cry elsewhere, Perona berated him for his lack of hospitality before listening with surprise as Mihawk revealed that he had actually seen Moria alive and well at the end of the war, though he could not definitively say whether or not he still lived.{{Qref|chap=592|ep=509|page=2-3|Perona learn of Moria's supposed death and is informed of its dubiousness by Mihawk.}}
</font>
 
  +
  +
Later, she helped an injured Zoro read the article about Luffy going to [[Marineford]] again by holding it up for him to read, though she complained about him taking a long time and her hands getting tired.{{Qref|chap=594|ep=512|page=16|Perona helps Zoro read the newspaper detailing Luffy's return to Marineford.}}
  +
  +
After accepting Zoro's request for training, Mihawk told Perona to tend to Zoro's wounds, prompting her to tell Mihawk to not order her around. Later, Perona asked Zoro about Luffy's message, so Zoro explained what it meant.{{Qref|chap=597|ep=515|page=5|Perona tends to Zoro's wounds and is informed of the meaning of Luffy's message.}}
  +
  +
===During the Timeskip===
  +
When Mihawk was teaching Zoro how to use [[Haki]], he forbade him from having any alcohol until he had mastered it, prompting Perona to laugh at him.{{Qref|chap=779|ep=720|page=3|Perona laughs at Zoro being unable to drink alcohol until he masters Haki.}}
  +
  +
{{Non-Canon}}
  +
  +
====3D2Y====
  +
While Luffy and [[Boa Hancock]] were fighting the [[World Pirates]], Perona traveled to their location with [[Dracule Mihawk]], where she helped them by inducing despair into [[Sebastian]] with her Negative Hollows, defeating him. She blushed when Luffy thanked her for the help before she lost sight of the pair.{{Qref|Special Episode [[3D2Y]], Perona aids Luffy and Hancock in the fight against the World Pirates.}}
  +
  +
{{Non-Canon|end}}
  +
  +
===Fish-Man Island Saga===
  +
====Return to Sabaody Arc====
  +
After Zoro's two year-long training, Perona took him back to [[Sabaody Archipelago]].{{Qref|name=Transporting Zoro}} As Luffy, Zoro, and Sanji made their way to the [[Thousand Sunny]], Perona aided them by fending off the Marines with her Negative Hollows before demanding to know why they were still waiting around when they should be preparing to leave. After scolding Zoro for his ungrateful attitude and brushing off Sanji's infatuation with her, Perona informed the trio that battleships were surrounding the island. Later, she assisted in the Straw Hats Pirates' escape by holding back the second Marine platoon with her Negative Hollows.{{Qref|chap=602|ep=522|page=4 & 13|Perona informs Luffy, Zoro, and Sanji of the Marine ships gathering around the archipelago and helps them escape.}}
  +
  +
====From the Decks of the World====
  +
  +
[[File:Perona Finds Kumashi.png|thumb|left|210px|Perona reunites with Kumashi.]]
  +
  +
Much later, she left Sabaody Archipelago and returned to the wandering Thriller Bark, where she discovered Kumashi's lifeless body, much to her joy.{{Qref|cover=658}}{{-}}
  +
  +
===Whole Cake Island Saga===
  +
====From the Decks of the World: The 500,000,000 Man Arc====
  +
Perona later returned to Kuraigana Island. After the events at [[Dressrosa]], Mihawk and Perona read the news of the Straw Hats' new bounties while tilling the land of Kuraigana Island with the humandrills.{{Qref|name=chap826}}
  +
===Wano Country Saga===
  +
{{Non-Canon}}
  +
====One Piece: Stampede====
  +
Perona, along with Mihawk, is one of the attendees at the [[Pirates Festival]]. Later, Perona used her Negative Hollow on [[Sentomaru]] and the Marines.{{Qref|movie=14}}
  +
{{Non-Canon|end}}
  +
  +
====Wano Country Arc====
  +
From reading a newspaper, Perona learned that Moria was alive and was annoyed that Mihawk did not inform her about it sooner. She then prepared to leave to go find him but not before thanking Mihawk for letting her stay with him.{{Qref|name=c925}}
  +
  +
===Final Saga===
  +
====Egghead Arc====
  +
Perona headed to [[Hachinosu]] and freed [[Koby]] from his imprisonment by the [[Blackbeard Pirates]], though she only did that to ask him help her in freeing Moria, who was also held prisoner.{{Qref|chap=1080|page=9|ep=|Perona frees Koby.}}
  +
  +
==Major Battles==
  +
*Perona vs. [[Usopp]]
  +
*Perona vs. [[Bartholomew Kuma]]
  +
*Perona vs. [[Sebastian]] (non-canon)
  +
*Perona vs. [[Marines]]
  +
  +
==Anime and Manga Differences==
  +
===Thriller Bark Arc===
  +
In the manga, when Usopp knocked her out, foam was coming out of her mouth;{{Qref|chap=466|page=12|Perona is foaming at the mouth after being knocked out by Usopp.}} this aspect of the scene was removed in the anime for unknown reasons.{{Qref|ep=361|Perona is not foaming at the mouth after being knocked out by Usopp.}}
  +
  +
===Straw Hat Separation Serial===
  +
  +
[[File:Zoro as Kumashi.png|thumb|210px|Perona dresses Zoro up as Kumashi.]]
  +
  +
In the anime version of Zoro's cover story, Perona displays more of a lonely childlike personality, mostly due to living by herself with no servants. She even tries to force Zoro to be her servant by using her Negative Hollows to make him swear to be her servant and dresses him up as [[Kumashi]].{{Qref|ep=456|Perona acts like a lonely child and tries to make Zoro her servant.}}
  +
  +
During the filler scenes on Kuraigana, although Perona was clearly in her ghost form, she had some interactions with physical objects despite supposedly being completely intangible.{{Qref|ep=456|Although Perona is in her ghost form, she raises dust upon falling on the ground.}}
  +
  +
In the anime, after Luffy's message is revealed, Perona went outside the castle with Mihawk and Zoro, where she scolded Zoro for not holding off on his training until his wounds were healed.{{Qref|ep=515|Perona scolds Zoro for continuing to train despite his injuries.}}
  +
  +
===Return to Sabaody Arc===
  +
When Perona is scolding Zoro at Sabaody, her hat bent in contact with the swordsman.{{Qref|ep=522|Perona's hat bends against Zoro while she is scolding him.}} This suggests that she was in her physical form rather than her ghost form; however, this does not occur in the manga.{{Qref|chap=602|page=4|Perona's hat does not bend upon making contact with Zoro.}} In the anime, Sanji is aroused by Perona's scent which also would mean she is in her physical form.
  +
  +
In the anime, Perona's interaction with Sanji is extended, where Sanji is beaten up and inflicted with depression by a Negative Hollow for freaking her out. Also, as she uses her ghosts to fend off the marines, Perona looks out to the sea and notes that she will have to find somebody else to play with, seemingly disappointed at Zoro's departure.{{Qref|ep=524|Perona attacks Sanji for freaking her out and expresses sadness at Zoro leaving.}}
  +
  +
==Early Concepts==
  +
An early concept of Perona from [[One Piece Green: Secret Pieces]] reveals that she was originally going to be a silent, princess-like girl.
  +
  +
{{Gallery|height=140|width=140|captionalign=center|position=center
  +
|Early Moria.png|Early Perona and her doll (Bottom left corner).
 
}}
 
}}
   
  +
==Translation and Dub Issues==
== External Links ==
 
  +
In the credits of [[One Piece: Stampede]], her name is spelled "Perhona".
* [[Wikipedia:Gothic Lolita|Gothic Lolita]] - Wikipedia article about the fashion style Perona wears.
 
  +
  +
==Merchandise==
  +
Perona features in the Portrait of Pirates series of figurines [http://mangafigures.wordpress.com/2011/01/23/ghost-princess-perona-excellent-model-one-piece-neo-dx-portraits-of-pirates-18-scale-pre-painted-pvc-figure/ in her pre-timeskip attire].
  +
  +
===Video Games===
  +
====Playable Appearances====
  +
*''[[One Piece: Pirate Warriors 2]]''
  +
*''[[One Piece: Pirate Warriors 3]]''
  +
*''[[One Py Berry Match]]''
  +
*''[[One Piece: Dance Battle]]''
  +
*[[One Piece: Burning Will]]
  +
*''[[One Piece: Burning Blood]]''
  +
*''[[One Piece Thousand Storm]]''
  +
*''[[One Piece Treasure Cruise]]''
  +
*''[[One Piece Bounty Rush]]''
  +
*''[[One Piece Bon! Bon! Journey!!]]''
  +
  +
====Support Appearances====
  +
*''[[One Piece: Gigant Battle]]''
  +
*''[[One Piece: Gigant Battle! 2 New World]]''
  +
*''[[One Piece: Great Pirate Colosseum]]''
  +
  +
====Enemy Appearances====
  +
*''[[One Piece Odyssey]]''
  +
  +
==Other Appearances==
  +
===Other Media===
  +
*Perona appears in the Zoro and Nami versions of the [[Hungry Days]] commercials.
  +
  +
==Trivia==
  +
[[File:Nami 4th Thriller Bark Outfit.png|thumb|100px|Perona's personal Jolly Roger on Nami's shirt.]]
  +
*Perona has a personal [[Jolly Roger]], which consists of a grinning skull with sharp teeth, wearing a crown resembling Perona's own. The skull has flames on the sides of its head, another pattern beneath it which matches the color of the flames, and a cross next to either side of the skull. [[Nami]] was seen wearing a top with this Jolly Roger on it, presumably having stolen it from Perona's room after disposing of the wedding dress Absalom forced her into.{{Qref|chap=475|ep=370|page=17|Nami wears a top with Perona's skull and crossbones.}}
  +
*In the 4th, 5th, 6th, and 7th [[Popularity Polls|fan polls]] of most popular characters, Perona was ranked 33rd, 21st, 23rd, and 25th, respectively.
  +
*Perona's obsession with cuteness and the style of clothing she wears both appear to be inspired by the [[wikipedia:Lolita fashion|Lolita subculture]].
  +
*Perona's favorite foods are bagel sandwiches and hot chocolate.{{Qref|name=vivre card}}
  +
{{-}}
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==References==
  +
{{References}}
   
 
==Site Navigation==
 
==Site Navigation==
{{Thriller Bark Navibox}}
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{{Thriller Bark Pirates Navibox}}
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{{Officers Navibox}}
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{{Paradise Characters Navibox}}
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{{Kuraigana Island Navibox}}
 
{{Devil Fruit Users Navibox}}
 
{{Devil Fruit Users Navibox}}
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{{West Blue Characters Navibox}}
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{{Straw Hat Pirates Allies Navibox}}
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[[ca:Perona]]
 
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[[fr:Perona]]
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[[id:Perona]]
 
[[it:Perona]]
 
[[it:Perona]]
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[[ja:ペローナ]]
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[[Category:Female Characters]]
 
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[[Category:Former Pirate Officers]]
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[[Category:Thriller Bark Pirates]]
 
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[[Category:Paramecia Devil Fruit Users]]
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[[Category:Mysterious Four]]
 

Latest revision as of 20:09, 19 March 2024

Featured Article Ahoy! This here is the 161st Featured Article.
"Perona" has been featured, meaning it was chosen as an article of interest.

"Ghost Princess" Perona[7] is the former commander of the Wild Zombies and Surprise Zombies of Thriller Bark prior to its collapse.[3] She was one of the Mysterious Four[2] and one of the major antagonists of the Thriller Bark Saga.

After the defeat of Gecko Moria's army and during the timeskip, she begrudgingly helped[8] and cooperated with Roronoa Zoro while in Dracule Mihawk's home on Kuraigana Island.[9]

Appearance

Perona is a short young woman with unusually big, round eyes, and she has long, light pink hair that she keeps tied into two pigtails with black and white flower hairpins. She wears a red crown with a black cross on her head, as well as red lipstick and a thick layer of black eyeliner. Her white long-sleeved three different black heart shapes shirt was short enough to expose her belly. She wears a red mini cape with a pink ribbon on front. Her black and white striped tights completely cover her legs, and she wears a mini-skirt and a red and gold belt with a heart belt buckle, as well as red buckled boots. She usually keeps her toes pointing inwards in a pigeon-toed manner and carried around a red parasol shaped like a cute version of a mini devil prior to her battle with Usopp.[3]

As a child, Perona carried the same parasol and wore the same crown and hairpins but wore her hair in much thicker pigtails and wore a simple, knee-length red dress with a white wave pattern and dots around its center, as well as smaller red boots. She also carried a ghost doll around on a leash and was accompanied by a bear named Kumae.[10]

When first meeting Hogback ten years prior to her encounter with the Straw Hat Pirates, Perona wore the same crown, but the rest of her outfit was not clearly visible, though she seemed to be wearing a similar dress.[11]

After the two-year timeskip, Perona has a more mature look than before, wearing a long black strapless dress with lavender frills, red high heeled boots and a big black top hat with white floral decor. She has a pink bat tattoo on her left bicep and a watch on her left wrist and wears pink nail polish on her fingernails. She still has her signature parasol, and also carries a stuffed animal resembling Kumashi with her. Additionally, she now wears her hair in multiple spiral braids on the back of her head.[12]

While reading about the Straw Hats' actions on Dressrosa and resulting new bounties, she wore a collared shirt with short, ruffled sleeves and a short dark tie hung loosely around her neck. She wore short frilly skirt with a line of crosses. She had short boots with bows on the front. She also had a large straw hat with a bow and large crown on top of it.[13]

Gallery

Main Series

Perona Pre Timeskip Portrait
A close up of Perona's face before the timeskip.
Perona Portrait
A close up of Perona's face after the timeskip.
Perona Post Timeskip Without Hat
Perona after the timeskip without her hat on.
Perona's Manga Color Scheme
Perona's color scheme in the manga before the timeskip.
Perona Post Timeskip Color Scheme
Perona's color scheme in the manga after the timeskip.
Perona Digitally Colored Manga After Timeskip
Perona's post-timeskip appearance in the Digitally Colored Manga.
Perona as a kid
Perona as a kid.
Perona 826
Perona's outfit during The 500,000,000 Man Arc.

Non-Canon

Perona Age 13
Perona at age 13.
Perona Wearing a Coat
Perona wearing a coat.
Perona Without Her Mini Cape
Perona without her mini cape.
Perona Boss Luffy Historical Special

Video Games

Perona Pirate Warriors 2
Perona DLC Pirate Warriors 2
Perona's Okuni DLC outfit in One Piece: Pirate Warriors 2.
Perona Wang Yuanji Pirate Warriors 2
Perona's Wang Yuanji DLC outfit in One Piece: Pirate Warriors 2.
Perona Thriller Bark Warriors 2
Perona's Thriller Bark DLC outfit in One Piece: Pirate Warriors 3.
Perona One Py Berry Match
Perona in One Py Berry Match.
Perona Super Grand Battle X
One Piece Burning Blood Swimsuit Perona (Artwork)
Artwork of Perona in her Swimsuit from One Piece: Burning Blood.
Perona Burning Will
Perona Thousand Storm
Perona before the timeskip in One Piece Thousand Storm.
Perona timeskip Thousand Storm
Perona after the timeskip in One Piece Thousand Storm.
Perona Summer Style Thousand Storm
Perona (Summer Style) in One Piece Thousand Storm.
Perona Ghost Princess Thousand Storm
Perona (Ghost Princess) in One Piece Thousand Storm.
Perona Wedding Thousand Storm
Perona (Wedding) in One Piece Thousand Storm.
Perona Valentine Thousand Storm
Perona (Valentine) in One Piece Thousand Storm.
Perona Bounty Rush
Perona Jumputi
Perona in Jumputi Heroes.
Beautiful Ghost Princess Perona Jumputi
"Beautiful Ghost Princess" Perona in Jumputi Heroes.

Other

Perona Anime Concept Art
Perona's concept art from the anime.
Kawaii Tokoro
Perona on the cover of the disc One Piece Nippon Judan! 47 Cruise CD.
One Piece novel HEROINES episode PERONA
Placole Perona
Perona's wedding dress from Placole Wedding Collaboration Dresses.
Perona Everyday
Perona in One Piece Everyday.

Personality

Perona's Shocked Face

Perona's face fault shocked at Ussop.

Perona has a very young, childlike and immature personality; frequently acting selfishly and on her own whims.[14][15] She dislikes people telling her what to do[16] and disobeying her will.[17] Despite this, she is normally loyal to Gecko Moria[18] and apparently sees him as a father-like role model, as she was crying to rejoin him after being sent to Kuraigana Island.[19] After Zoro appeared on Kuraigana as well, she tended to his wounds, displaying a willingness to befriend even those she formerly knew as a foe and demonstrating that she can be kind to others, despite her normally cruel demeanor.[8]

She is very fond of cute things and will only allow someone to work for her if they are cute.[20] Her dream is to create a kingdom where all the cute things in the world have been turned into zombie animals that obey only her.[21] She dislikes Kumashi because, although he is cute in appearance, his voice is not, and thus she prefers him to just be quiet.[22] Her desire for cute things gives her a slightly childish outlook on some issues.[20] She is somewhat playful during combat, threatening to crush Usopp's heart when she was in her ghost form, which nearly caused Usopp to die from fear, and shortly afterward merely laughing it off when revealing that she could not do so.[23]

Perona Screaming In Fear

Perona screams in fear at being covered in cockroaches.

Perona has katsaridaphobia, a fear of cockroaches, to the point of screaming and crying when Usopp covered her with fake ones.[24] She is also somewhat naive, having watched Usopp seemingly lift a 10-ton hammer without questioning whether or not his feat was genuine, and shortly afterward passing out from fear when he attacked her with it, despite not actually being harmed.[25]

She is very confident in her Devil Fruit powers being able to defeat anyone.[26] Even when confronted by the powerful and dreaded Warlord of the Sea Bartholomew Kuma, after initially being frozen with fear, Perona quickly regained her confidence and proclaimed that she would defeat him.[27] She is also quite stubborn about using them in combat, having continued to use her Negative Hollows against Usopp even after they proved useless due to his negativity because she believed that his immunity was a fluke.[28]

After the timeskip, she has become more mature, although she still carries a teddy bear around, indicating that she is still obsessed with cuteness to some degree.[12] She displayed a grateful side, as she tearfully thanked Dracule Mihawk for letting her stay on his island.[29]

Like many others, Perona has her own unique laugh, which uses the word Horo (e.g., "Horo-horo-horo-horo-horo!"); this is based on the name of her Devil Fruit, the Horo Horo no Mi.[30]

Relationships

Thriller Bark Pirates

Perona followed her master, Gecko Moria, for at least ten years.[11] Her loyalty to him is somewhat complex: when she sensed defeat, she attempted to leave and abandon him[15] while noting that she had only stayed with him for fun,[31] but tried to take Bartholomew Kuma down as a final gift for Moria.[32] Despite not clearly showing it, she is said to love Moria like a father since he found her and raised her as a child, and when the newspaper falsely reported that Moria died in the Whitebeard War, Perona was deeply saddened.[33][34] When she learned that Moria is alive two years later, Perona is overjoyed, and she quickly departed Mihawk's castle to go find him.[35]

Perona had a mixed relationship with her teddy bear servant, Kumashi. While she considered him to be cute in appearance, she forbade him from speaking because his voice is not cute.[22] However, she was devastated when Usopp purified Kumashi.[36]

Straw Hat Pirates

Perona Scolds Zoro

Perona scolds Zoro.

While on Thriller Bark, Perona became enemies with one particular member of the Straw Hat Pirates, Usopp, due to the latter becoming the first person to ever resist the effects of her Negative Hollows[37] and later defeating her in a humiliating manner.[25] However, she also cannot help but pity Usopp who lives his life in negativity and attempts to cheer him up, though this may be for her own benefit, as she cannot make him negative unless he was already slightly positive.[38][39] She looked down on Nami when the latter confronted her later, claiming that she could easily defeat Nami with her Mini Hollows because Usopp was her only natural enemy.[40]

After encountering Zoro on Kuraigana Island, she was shocked to see him but decided to treat his wounds as he was the only other person there at the time.[8] Despite their initial quarrels,[41] the two developed a relationship of mutual trust by living on the island for two years alongside Dracule Mihawk.[42] She also willingly brought Zoro back to Sabaody Archipelago[43] and later held back the Marines in order to allow the Straw Hats to escape.[44]

Dracule Mihawk

After being sent to Kuraigana Island by Kuma's abilities, Perona decided to take residence in the castle where the at the time Warlord Mihawk lived.[45] Mihawk seemingly agreed or simply did not mind letting her stay with him, though he did not display any hospitality toward her beyond that.[46]

It was shown in From the Decks of the World: The 500,000,000 Man Arc that even after going back to Thriller Bark to retrieve Kumashi, she returned to Kuraigana Island and was seen helping him with his work in tilling the land, suggesting a good status between them.[47] When she learned of her old master's survival, she was upset that Mihawk did not tell her and nearly left as he told her to be safe while she tearfully thanked him for letting her stay with him.[35]

Abilities and Powers

Being formerly one of the Mysterious Four, Perona had a number of zombies under her command prior to her defection from Thriller Bark.[3] Her ex-crewmate, Doctor Hogback, acknowledges her value under Moria's employment.[48]

Perona is physically strong, as she was seen carrying Kumashi's body, which is bigger than she is, while levitating with her Devil Fruit powers.

Perona is able to cook, having cooked several times during the two years while she was living with Mihawk.[35]

Medical Knowledge

While she seems to recognize that bandages can help a person who is wounded, she seems to lack expertise in medicine. During the process of treating Zoro's wounds, she covered him from head to toe in an overabundance of bandages the first time she found him.[8] However, despite this fault, she has common sense or enough knowledge of medical skills, as Zoro recovered from his injuries.[49]

Navigation Knowledge

She is a capable navigator, demonstrated when she brought Zoro to Sabaody Archipelago all the way from Kuraigana Island.[43] She later traveled to Thriller Bark and returned back to Kuraigana Island all by herself. After learning Moria was alive, she traveled to all the way to Hachinosu from Kuraigana Island, going from Paradise to the New World.

Devil Fruit

Further information: Horo Horo no Mi
Horo Horo no Mi Power

Perona's negative ghosts mocking her enemies.

Perona ate the Horo Horo no Mi, a Paramecia-type Devil Fruit that allows her to create and control ghosts. The ghosts have the power to drain the morale, self-esteem, and will to live of anyone whom they pass through, described as hollowing out the victim's spirit.[50][51] She can also create smaller ghosts that explode with enough force to damage stone walls[52] and larger ghosts that produce an even more powerful blast.[53]

Her ghosts can also be used to form a "Ghost Network" for surveillance. This allows her to spy on enemies.[54][55] The ghosts can also duplicate themselves into even more ghosts.[56]

In addition to creating ghosts, she can make her own ghost leave her body. Her disembodied ghost is completely intangible; she can pass through solid objects and cannot be hit by enemy attacks, although she cannot damage anything or attack her opponents physically.[57][58] Her ghost also has the power to levitate and can change the size of her spectral body.[59] However, while she is doing this, her true body remains unconscious and vulnerable.[60]

History

Past

When she was a child, Perona owned a pet bear named Kumae.[10] At some point in her childhood, she met the pirate Gecko Moria who took her in and raised her and she eventually saw him as a father figure.[34]

Hogback Meets Moria, Absalom and Perona

Perona witnesses the meeting between Moria and Hogback.

Over 10 years ago, she joined Moria's crew merely for fun.[31] She and Absalom were present when Moria met and recruited the genius doctor, Hogback.[11]

Five years ago, she was sleeping while Brook was causing trouble in Thriller Bark, which would later cause her to not know what he looked like when he returned.[61] When Kumashi tried to wake her up, she simply told him to shut up and went back to sleep.[62] Three and a half years ago, Perona came to Moria with news that Bartholomew Kuma was elected the new Warlord of the Sea, Moria brushed it off finding his moniker funny but Perona told him that he must be a strong figure to be chosen as a Warlord.[63]

Thriller Bark Saga

Thriller Bark Arc

Perona used her Ghost Network to scout out the Thousand Sunny[1] and later reported the crew's bounties to Hogback and Absalom.[64]

In the Dead Forest, a group of Negative Hollows encountered some of the Straw Hats and, after evading capture, demoralized Monkey D. Luffy, Franky, and Roronoa Zoro.[65] As the Night Attack began, Perona recalled her ghosts and rallied her Wild Zombies and Surprise Zombies.[66] When she left to oversee the looting of the Thousand Sunny, she ordered Kumashi to meet the Risky Brothers, who were carrying Nami, Usopp, and Tony Tony Chopper in coffins, and take them to the Dance Hall.[67]

On the Thousand Sunny, Perona was frustrated at the lack of treasure and had the food taken from the ship. Hildon reported to Perona that Moria was gathering the other members of the Mysterious Four because he had captured Luffy and would be debuting Zombie 900, much to her surprise. Soon afterward, Perona returned to her room, where she found Absalom speaking with Kumashi about the escape of Nami, Usopp, and Chopper. After scolding Kumashi for trying to speak, Perona informed Absalom that Moria was calling them together and explained why he was doing so.[68] In the Mast Mansion, the Mysterious Four discussed Luffy and the Straw Hats, and Perona reported that Nami, Usopp, and Chopper had escaped before yelling at Kumashi when he tried to correct her. When Luffy broke free from his cage, Perona quickly stopped him with her Negative Hollow before watching on as Moria removed Luffy's shadow.[69] After hearing Nami, Usopp, and Chopper talking inside Kumashi and believing that it was Kumashi himself doing so, Perona proclaimed that her dream of creating a kingdom full of cute subordinates was closer to fruition with Luffy's shadow extracted. When two Spider Mice appeared and reported that Tararan's shadow had been extracted, Hogback noted that it could only have been the "Humming Swordsman", a name which Perona did not recognize. Soon afterward, Perona accompanied the rest of the Mysterious Four to the enormous freezer containing the corpse of Oars.[70] Perona watched as Moria used Luffy's shadow to revive Oars's corpse. When Oars awoke, Nami, Usopp, and Chopper screamed and fell from Kumashi's back, prompting Perona to yell at Kumashi for hiding the pirates.[71] Perona continued to chastise Kumashi for hiding the pirates without letting him respond.[72]

After reporting Oars's current activities to Moria, Perona learned that the Straw Hats had learned of the zombies' weakness to salt and were approaching them, with the General Zombies preoccupied with attending Absalom's wedding, but dismissed Hogback's worry over the zombies' weakness being known and retired to her room before attacking the Straw Hats with her ghosts, planning to weaken them enough to be taken by Soldier Zombies. However, Perona's attack was interrupted by Oars crashing down through the mansion.[73] After Franky rebuilt the destroyed bridge, he, Zoro, Sanji, and Usopp charged into Perona's room, where Perona confronted them. After explaining the powers granted by her Devil Fruit, Perona sent a Negative Hollow through each of the Straw Hats, but Usopp was unaffected and purified several of her zombies before declaring that he was always negative, shocking Perona, who tried to cheer him up alongside her zombies before preparing to battle him.[38]

Perona vs

Perona vs. Usopp.

Believing Usopp's immunity to have been a fluke, Perona sent four Negative Hollows through him, but instead became depressed herself alongside her ghosts, to the shock of her zombies.[74] Soon afterward, Perona fled from Usopp while noting that she could defeat the other Straw Hats if it were not for him. After Kumashi distracted Usopp by attacking him, Perona projected her spirit from her body after hiding and tricked Usopp into believing she was flying while proclaiming that she had come up with a plan to defeat him.[39] Declaring that she had simply been caught up in the moment before, Perona attempted to cheer Usopp up once more before appearing behind him in a giant form. After dodging Usopp's Ageha Ryusei, Perona reverted to her normal size before placing her hands inside of Usopp's chest and pretending that she was going to crush his heart, only to reveal that she could not do such a thing. Usopp attacked her with Rokuren Mamushi Boshi, but Perona demonstrated her intangibility by letting the attack go through her and moving through the floor. When Usopp pointed out that she could not touch him either and demanded to know how she could attack him, Perona summoned some Mini Hollows and demonstrated their explosive power on a nearby wall before attacking Usopp with one as well. After tricking Usopp into running into a pillar, Perona covered his body with Mini Hollows and detonated all of them before gleefully watching Kumashi brutalize Usopp. However, Perona was shocked to see Usopp, who had donned his Sogeking mask, purify Kumashi with a salt ball before fleeing once more. Wounding Usopp with a barrage of Mini Hollow blasts once more, Perona decided to hand him over to Moria, but was surprised when Usopp figured out her trick and used Atlas Suisei to expose the room containing her true body.[75]

Usopp Defeats Perona

Usopp defeats Perona.

After Usopp deduced how she left her body by becoming a ghost in order to fight him, Perona revealed that she was a fully conscious Hollow instead of merely being a ghost controlled by her body, prompting Usopp to seemingly attack her body in an attempt to destroy it. After the attack apparently missed, a panicked Perona engulfed his body with a Toku Hollow before returning to her own body. Perona detonated the Toku Hollow but was surprised to see the explosion had disappeared as Usopp revealed he had absorbed it with an Impact Dial. After learning that Usopp had actually shot a Torimochi Boshi at her bed in order to trap her body in one place, Perona witnessed Usopp defeat Hippo Gentleman with a blast from his Impact Dial before covering Perona herself in fake cockroaches with Kurobikari Boshi. Crying and screaming hysterically at the sight of cockroaches, a shocked Perona saw Usopp seemingly lifting a 10-ton hammer and begged him to spare her before passing out in fright as the fake hammer burst apart on impact with her. Usopp celebrated his victory over her as Perona collapsed while foaming at the mouth.[76]

Later, Perona's zombies found Perona still unconscious but mostly unharmed as Oars rampaged through Thriller Bark.[77] Soon afterward, Perona regained consciousness and had to be calmed down by her zombies when she acted like Usopp was still attacking her. Upon seeing the damage done to the mansion by Oars, Perona wondered if the Straw Hats had done this and was shocked to learn that Oars had completely wiped out the General Zombies as well, which caused her to question if they could control him at all. However, the zombies' explanation of what was happening was interrupted by Oars punching them through one of the nearby walls while searching for Nami; witnessing this, Perona was convinced that Oars and the Straw Hats were going to completely destroy Thriller Bark and ordered her remaining subordinates to gather all the treasure and some supplies so they could leave on the Thousand Sunny.[78]

Perona Encounters Kuma

Perona meets Kuma.

Observing the battle between the Straw Hats and Oars from the Thousand Sunny, Perona admitted that she only joined Moria for fun and would not stay with him if her life were in danger before telling her subordinates that they would be leaving without delay.[79] When Nami confronted her over her taking the treasure and the Thousand Sunny, Perona declared that she was planning to leave Thriller Bark on the Thousand Sunny and prepared to battle while proclaiming that her Mini Hollows should be enough to defeat Nami. However, she was interrupted by the appearance of Bartholomew Kuma, whom Perona was terrified at the sight of due to knowing his reputation as a cruel pirate and a Warlord of the Sea. When Kuma asked her where she would like to go on a trip, Perona described a dark, damp island with an old castle filled with malice before realizing that she had gotten caught up in his pace. Resolving to take down Kuma as a parting gift for Moria due to his shadow likely being strong, Perona prepared to attack him with a combination of Negative Hollows and a Toku Hollow, but Kuma suddenly made her disappear before her attack could connect, to the confusion of those watching.[80]

When Hogback and Absalom escaped Thriller Bark later, they left without Perona while noting they could not find her anywhere.[81]

Summit War Saga

Amazon Lily Arc

Perona Finds Zoro

Perona encounters Zoro.

Kuma honored Perona's request, having sent her to Kuraigana Island, an island filled with dark and spooky ruins, in addition to the malicious castle. Although she had taken up residence in the castle, Perona was not happy because she had no servants or cute animals. As she lamented her situation while wishing she was back on Thriller Bark, Perona noticed a disturbance in the forest and discovered that Zoro had been sent to the island as well. Initially terrified by the sight of one of the Straw Hats, Perona eventually decided to save him since he was the only other person there at the time and brought him to the castle, where she treated his wounds by heavily bandaging him. When Zoro regained consciousness, he found out that she had taken his swords away to prevent him from attacking her.[82]

Straw Hat's Separation Serial

Eventually, Perona returned Zoro's swords and followed him through the ruins of the island, indicating that they had developed some level of trust by then.[83]

Post War Arc

Mihawk at Home

Perona and Mihawk discuss Moria's fate.

After the Summit War of Marineford, the owner of the castle that Perona was inhabiting, Dracule Mihawk, returned to Kuraigana Island, and Perona began sobbing at the news of Moria's death in the newspaper. When Mihawk told her to cry elsewhere, Perona berated him for his lack of hospitality before listening with surprise as Mihawk revealed that he had actually seen Moria alive and well at the end of the war, though he could not definitively say whether or not he still lived.[84]

Later, she helped an injured Zoro read the article about Luffy going to Marineford again by holding it up for him to read, though she complained about him taking a long time and her hands getting tired.[85]

After accepting Zoro's request for training, Mihawk told Perona to tend to Zoro's wounds, prompting her to tell Mihawk to not order her around. Later, Perona asked Zoro about Luffy's message, so Zoro explained what it meant.[86]

During the Timeskip

When Mihawk was teaching Zoro how to use Haki, he forbade him from having any alcohol until he had mastered it, prompting Perona to laugh at him.[87]


The following events are Non-Canon and therefore not considered part of the Canon story.


3D2Y

While Luffy and Boa Hancock were fighting the World Pirates, Perona traveled to their location with Dracule Mihawk, where she helped them by inducing despair into Sebastian with her Negative Hollows, defeating him. She blushed when Luffy thanked her for the help before she lost sight of the pair.[88]


Concludes non-canon section.


Fish-Man Island Saga

Return to Sabaody Arc

After Zoro's two year-long training, Perona took him back to Sabaody Archipelago.[43] As Luffy, Zoro, and Sanji made their way to the Thousand Sunny, Perona aided them by fending off the Marines with her Negative Hollows before demanding to know why they were still waiting around when they should be preparing to leave. After scolding Zoro for his ungrateful attitude and brushing off Sanji's infatuation with her, Perona informed the trio that battleships were surrounding the island. Later, she assisted in the Straw Hats Pirates' escape by holding back the second Marine platoon with her Negative Hollows.[89]

From the Decks of the World

Perona Finds Kumashi

Perona reunites with Kumashi.

Much later, she left Sabaody Archipelago and returned to the wandering Thriller Bark, where she discovered Kumashi's lifeless body, much to her joy.[90]

Whole Cake Island Saga

From the Decks of the World: The 500,000,000 Man Arc

Perona later returned to Kuraigana Island. After the events at Dressrosa, Mihawk and Perona read the news of the Straw Hats' new bounties while tilling the land of Kuraigana Island with the humandrills.[13]

Wano Country Saga


The following events are Non-Canon and therefore not considered part of the Canon story.


One Piece: Stampede

Perona, along with Mihawk, is one of the attendees at the Pirates Festival. Later, Perona used her Negative Hollow on Sentomaru and the Marines.[91]


Concludes non-canon section.


Wano Country Arc

From reading a newspaper, Perona learned that Moria was alive and was annoyed that Mihawk did not inform her about it sooner. She then prepared to leave to go find him but not before thanking Mihawk for letting her stay with him.[35]

Final Saga

Egghead Arc

Perona headed to Hachinosu and freed Koby from his imprisonment by the Blackbeard Pirates, though she only did that to ask him help her in freeing Moria, who was also held prisoner.[92]

Major Battles

Anime and Manga Differences

Thriller Bark Arc

In the manga, when Usopp knocked her out, foam was coming out of her mouth;[93] this aspect of the scene was removed in the anime for unknown reasons.[94]

Straw Hat Separation Serial

Zoro as Kumashi

Perona dresses Zoro up as Kumashi.

In the anime version of Zoro's cover story, Perona displays more of a lonely childlike personality, mostly due to living by herself with no servants. She even tries to force Zoro to be her servant by using her Negative Hollows to make him swear to be her servant and dresses him up as Kumashi.[95]

During the filler scenes on Kuraigana, although Perona was clearly in her ghost form, she had some interactions with physical objects despite supposedly being completely intangible.[96]

In the anime, after Luffy's message is revealed, Perona went outside the castle with Mihawk and Zoro, where she scolded Zoro for not holding off on his training until his wounds were healed.[97]

Return to Sabaody Arc

When Perona is scolding Zoro at Sabaody, her hat bent in contact with the swordsman.[98] This suggests that she was in her physical form rather than her ghost form; however, this does not occur in the manga.[99] In the anime, Sanji is aroused by Perona's scent which also would mean she is in her physical form.

In the anime, Perona's interaction with Sanji is extended, where Sanji is beaten up and inflicted with depression by a Negative Hollow for freaking her out. Also, as she uses her ghosts to fend off the marines, Perona looks out to the sea and notes that she will have to find somebody else to play with, seemingly disappointed at Zoro's departure.[100]

Early Concepts

An early concept of Perona from One Piece Green: Secret Pieces reveals that she was originally going to be a silent, princess-like girl.

Early Moria
Early Perona and her doll (Bottom left corner).

Translation and Dub Issues

In the credits of One Piece: Stampede, her name is spelled "Perhona".

Merchandise

Perona features in the Portrait of Pirates series of figurines in her pre-timeskip attire.

Video Games

Playable Appearances

Support Appearances

Enemy Appearances

Other Appearances

Other Media

  • Perona appears in the Zoro and Nami versions of the Hungry Days commercials.

Trivia

Nami 4th Thriller Bark Outfit

Perona's personal Jolly Roger on Nami's shirt.

  • Perona has a personal Jolly Roger, which consists of a grinning skull with sharp teeth, wearing a crown resembling Perona's own. The skull has flames on the sides of its head, another pattern beneath it which matches the color of the flames, and a cross next to either side of the skull. Nami was seen wearing a top with this Jolly Roger on it, presumably having stolen it from Perona's room after disposing of the wedding dress Absalom forced her into.[101]
  • In the 4th, 5th, 6th, and 7th fan polls of most popular characters, Perona was ranked 33rd, 21st, 23rd, and 25th, respectively.
  • Perona's obsession with cuteness and the style of clothing she wears both appear to be inspired by the Lolita subculture.
  • Perona's favorite foods are bagel sandwiches and hot chocolate.[4]


References

  1. 1.0 1.1 One Piece Manga and Anime — Vol. 46 Chapter 443 (p. 15) and Episode 338, Perona appears as a ghost before the Straw Hats.
  2. 2.0 2.1 One Piece Manga and Anime — Vol. 47 Chapter 455 (p. 11-13) and Episode 349, Perona is identified as one of the Mysterious Four.
  3. 3.0 3.1 3.2 3.3 3.4 One Piece Manga and Anime — Vol. 46 Chapter 449 (p. 18) and Episode 343, Perona is formally introduced.
  4. 4.0 4.1 4.2 Vivre Card - One Piece Visual Dictionary (Card #0473), Information about Perona is revealed.
  5. 5.0 5.1 SBS One Piece Manga — Vol. 82 (p. 102), Perona's pre and post timeskip ages and height are revealed.
  6. One Piece Blue Deep: Characters World (p. 157), Perona's birthday is revealed.
  7. One Piece Manga — Vol. 92 Chapter 925 (p. 2), Perona's name is romanized.
  8. 8.0 8.1 8.2 8.3 One Piece Manga and Anime — Vol. 54 Chapter 524 (p. 12) and Episode 421, Perona treats Zoro's injuries after finding him.
  9. One Piece Manga and Anime — Vol. 61 Chapter 597 (p. 5) and Episode 515, Perona prepares to help Zoro heal so he can begin his training.
  10. 10.0 10.1 SBS One Piece Manga — Vol. 66 (p. 182), Perona is drawn as a child.
  11. 11.0 11.1 11.2 One Piece Manga and Anime — Vol. 48 Chapter 468 (p. 9) and Episode 363, Perona accompanies Moria to Hogback's recruitment.
  12. 12.0 12.1 One Piece Manga and Anime — Vol. 61 Chapter 602 (p. 4) and Episode 522, Perona appears on the Sabaody Archipelago after the timeskip.
  13. 13.0 13.1 One Piece Manga — Vol. 82 Chapter 826, cover story: From the Decks of the World: The 500,000,000 Man Arc Vol. 17.
  14. One Piece Manga and Anime — Vol. 47 Chapter 451 (p. 6) and Episode 345, Perona decides to oversee the looting of the Thousand Sunny herself.
  15. 15.0 15.1 One Piece Manga and Anime — Vol. 49 Chapter 471 (p. 6) and Episode 366, Perona decides to flee from Thriller Bark with all the treasure.
  16. One Piece Manga and Anime — Vol. 61 Chapter 597 (p. 5) and Episode 515, Perona angrily tells Mihawk to not order her around.
  17. One Piece Manga and Anime — Vol. 47 Chapter 454 (p. 9) and Episode 348, Perona is irritated by Kumashi continuing to speak after she told him not to.
  18. One Piece Manga and Anime — Vol. 48 Chapter 460 (p. 11) and Episode 354, Perona declares that she will take out the Straw Hats by herself after learning that they have discovered the zombies' weakness to salt.
  19. One Piece Manga and Anime — Vol. 54 Chapter 524 (p. 10) and Episode 421, Perona expresses her desire to return to Thriller Bark.
  20. 20.0 20.1 One Piece Manga and Anime — Vol. 46 Chapter 449 (p. 12) and Episode 343, Perona declares that only cute beings may work for her.
  21. One Piece Manga and Anime — Vol. 47 Chapter 456 (p. 10) and Episode 350, Perona reveals her dream of creating a kingdom of cute subordinates.
  22. 22.0 22.1 One Piece Manga and Anime — Vol. 46 Chapter 449 (p. 12) and Episode 343, Perona is annoyed by Kumashi's deep voice contrasting with his cute appearance.
  23. One Piece Manga and Anime — Vol. 48 Chapter 465 (p. 6-8) and Episode 360, Perona tricks Usopp into believing that she will crush his heart.
  24. One Piece Manga and Anime — Vol. 48 Chapter 466 (p. 9) and Episode 361, Perona admits her fear of cockroaches upon being covered in fake ones.
  25. 25.0 25.1 One Piece Manga and Anime — Vol. 48 Chapter 466 (p. 10-12) and Episode 361, Perona passes out in fear after being scared by Usopp's tricks.
  26. One Piece Manga and Anime — Vol. 49 Chapter 473 (p. 15) and Episode 368, Perona claims that there is no one whom her powers cannot defeat.
  27. One Piece Manga and Anime — Vol. 49 Chapter 473 (p. 12-15) and Episode 368, Perona is initially frightened by Kuma, but quickly regains her resolve.
  28. One Piece Manga and Anime — Vol. 48 Chapter 462 (p. 2-3) and Episode 357, Perona stubbornly continues to use her Negative Hollows against Usopp.
  29. One Piece Manga — Vol. 92 Chapter 925.
  30. One Piece Manga and Anime — Vol. 46 Chapter 449 (p. 12 & 18) and Episode 343, Perona laughs while preparing to join the assault on the Straw Hats.
  31. 31.0 31.1 One Piece Manga and Anime — Vol. 49 Chapter 472 (p. 18) and Episode 367, Perona admits that she had only worked with Moria for fun.
  32. One Piece Manga and Anime — Vol. 49 Chapter 473 (p. 15) and Episode 368, Perona decides to take down Kuma as a parting gift for Moria.
  33. One Piece Manga and Anime — Vol. 60 Chapter 592 (p. 2) and Episode 509, Perona grieves over the false report of Moria's death.
  34. 34.0 34.1 SBS One Piece Manga — Vol. 82 (p. 102), Perona's past with Moria is explained.
  35. 35.0 35.1 35.2 35.3 One Piece Manga and Anime — Vol. 92 Chapter 925 (p. 2-3) and Episode 917, Perona learns Moria is alive and leaves Mihawk.
  36. One Piece Manga and Anime — Vol. 48 Chapter 465 (p. 16) and Episode 360, Perona is shocked and angered at the sight of Usopp purifying Kumashi.
  37. One Piece Manga and Anime — Vol. 48 Chapter 462 (p. 2-3) and Episode 357, Usopp resists the effects of four Negative Hollows at once, causing the ghosts and Perona herself to become depressed instead.
  38. 38.0 38.1 One Piece Manga and Anime — Vol. 48 Chapter 461 (p. 11-18) and Episode 356, Perona confronts several of the Straw Hats, but is shocked to learn Usopp is immune to her Negative Hollows.
  39. 39.0 39.1 One Piece Manga and Anime — Vol. 48 Chapter 464 (p. 16-19) and Episode 359, Perona flees from Usopp before exiting her body in order to begin her counterattack.
  40. One Piece Manga and Anime — Vol. 49 Chapter 473 (p. 11) and Episode 368, Perona talks down to Nami.
  41. One Piece Manga and Anime — Vol. 54 Chapter 524 (p. 12) and Episode 421, Perona and Zoro argue over Perona taking away Zoro's swords.
  42. One Piece Manga and Anime — Vol. 61 Chapter 602 (p. 4) and Episode 522, Perona interacts with Zoro after bringing him to Sabaody Archipelago.
  43. 43.0 43.1 43.2 One Piece Manga and Anime — Vol. 61 Chapter 602 (p. 4) and Episode 522, Perona reveals that she brought Zoro back to Sabaody Archipelago.
  44. One Piece Manga and Anime — Vol. 61 Chapter 602 (p. 13) and Episode 522, Perona uses her Negative Hollows to demoralize a Marine unit.
  45. One Piece Manga and Anime — Vol. 54 Chapter 524 (p. 10) and Episode 421, Perona begins inhabiting Mihawk's castle.
  46. One Piece Manga and Anime — Vol. 60 Chapter 592 (p. 2) and Episode 509, Mihawk allows Perona to stay in his castle, but does not display any more hospitality than that toward her.
  47. One Piece Manga — Vol. 82 Chapter 826.
  48. One Piece Manga and Anime — Vol. 48 Chapter 460 (p. 11) and Episode 355, Hogback admits that they are lucky to have Perona on their side.
  49. One Piece Manga and Anime — Vol. 61 Chapter 597 (p. 5) and Episode 515, Perona is entrusted with ensuring Zoro's wounds heal.
  50. One Piece Manga and Anime — Vol. 48 Chapter 461 (p. 13) and Episode 356, Perona reveals the name of her Devil Fruit and explains its power.
  51. One Piece Manga and Anime — Vol. 46 Chapter 448 (p. 9-10) and Episode 342, Franky, Luffy, and Zoro are made negative by the ghosts passing through them.
  52. One Piece Manga and Anime — Vol. 48 Chapter 465 (p. 10-11) and Episode 360, Perona demonstrates the power of her Mini Hollows
  53. One Piece Manga and Anime — Vol. 48 Chapter 466 (p. 5-6 & *) and Episode 361, Perona uses the Toku Hollow, and Usopp absorbs the blast with an Impact Dial and uses it to defeat Hippo Gentleman.
  54. One Piece Manga and Anime — Vol. 47 Chapter 453 (p. 13) and Episode 347, Tararan identifies Perona's surveillance system as her "ghost network".
  55. One Piece Manga and Anime — Vol. 46 Chapter 443 (p. 16) and Episode 338, Brook notes the Thousand Sunny is already under surveillance by Perona's ghosts.
  56. One Piece Manga and Anime — Vol. 46 Chapter 448 (p. 8) and Episode 342, One of Perona's ghosts splits into three ghosts.
  57. One Piece Manga and Anime — Vol. 48 Chapter 465 (p. 9) and Episode 360, Perona demonstrates that all attacks will go through her astral projection.
  58. One Piece Manga and Anime — Vol. 48 Chapter 465 (p. 6-8) and Episode 360, Perona demonstrates her ability to pass through solid matter, but reveals that she cannot physically interact with it.
  59. One Piece Manga and Anime — Vol. 48 Chapter 465 (p. 4-5) and Episode 360, Perona changes the size of her astral projection.
  60. One Piece Manga and Anime — Vol. 48 Chapter 466 (p. 4) and Episode 361, Perona panics over Usopp attacking her real body.
  61. One Piece Manga and Anime — Vol. 47 Chapter 456 (p. 12) and Episode 350, Perona reveals that she does not know what Brook looks like.
  62. One Piece Manga and Anime — Vol. 47 Chapter 458 (p. 11) and Episode 352, Perona sleeps through the crisis caused by Brook on Thriller Bark.
  63. One Piece Manga — Vol. 108 Chapter 1100 (p. 16), Perona tells Moria that a new Warlord was elected.
  64. One Piece Manga and Anime — Vol. 46 Chapter 447 (p. 7) and Episode 341, Perona reports the bounties of the Straw Hats to Absalom and Hogback via one of her ghosts.
  65. One Piece Manga and Anime — Vol. 46 Chapter 448 (p. 8-10) and Episode 342, Perona's ghosts demoralize some of the Straw Hats in the forest.
  66. One Piece Manga and Anime — Vol. 46 Chapter 449 (p. 12 & 18) and Episode 343, Perona returns to her bodies and rallies her zombies.
  67. One Piece Manga and Anime — Vol. 47 Chapter 451 (p. 6) and Episode 345, Perona orders Kumashi to assist the Risky Brothers.
  68. One Piece Manga and Anime — Vol. 47 Chapter 454 (p. 7-9) and Episode 348, Perona learns of Moria summoning the other members of the Mysterious Four and informs Absalom of it as well.
  69. One Piece Manga and Anime — Vol. 47 Chapter 455 (p. 11-19) and Episode 349, Perona meets with the rest of the Mysterious Four and incapacitates Luffy.
  70. One Piece Manga and Anime — Vol. 47 Chapter 456 (p. 8, 10-12 & 17-19) and Episode 350, Perona learns of Brook's return and enters the freezer containing Oars's corpse.
  71. One Piece Manga and Anime — Vol. 47 Chapter 457 (p. 7-9) and Episode 351, Perona witnesses Oars's awakening and learns of the Straw Hats hiding inside of Kumashi.
  72. One Piece Manga and Anime — Vol. 47 Chapter 458 (p. 4) and Episode 352, Perona berates Kumashi for hiding the Straw Hats.
  73. One Piece Manga and Anime — Vol. 48 Chapter 460 (p. 8, 11 & 15-17) and Episode 355, Perona attempts to weaken the Straw Hats with her Negative Hollows so they cannot purify any zombies but is interrupted by Oars.
  74. One Piece Manga and Anime — Vol. 48 Chapter 462 (p. 2-3) and Episode 357, Perona becomes negative after attempting to overwhelm Usopp with four Negative Hollows at once.
  75. One Piece Manga and Anime — Vol. 48 Chapter 465 (p. 2-19) and Episode 360, Perona battles Usopp and initially overwhelms him, but has her true body exposed.
  76. One Piece Manga and Anime — Vol. 48 Chapter 466 (p. 2-12) and Episode 361, Usopp outmaneuvers and defeats Perona.
  77. One Piece Manga and Anime — Vol. 48 Chapter 469 (p. 17) and Episode 364, An unconscious Perona is found by her zombies.
  78. One Piece Manga and Anime — Vol. 49 Chapter 471 (p. 4-6) and Episode 366, Perona decides to leave Thriller Bark after witnessing the destruction caused by Oars.
  79. One Piece Manga and Anime — Vol. 49 Chapter 472 (p. 18-19) and Episode 367, Perona notes why she is leaving Thriller Bark while watching the battle against Oars.
  80. One Piece Manga and Anime — Vol. 49 Chapter 473 (p. 10-16) and Episode 368, Perona confronts Kuma, but is effortlessly defeated.
  81. One Piece Manga and Anime — Vol. 50 Chapter 486 (p. 2) and Episode 378, Hogback and Absalom could not find Perona.
  82. One Piece Manga and Anime — Vol. 54 Chapter 524 (p. 10-12) and Episode 421, Perona laments her situation and discovers Zoro has been sent to Kuraigana Island as well.
  83. One Piece Manga and Anime — Vol. 57 Chapter 559, cover story: Straw Hat's Separation Serial #8 Vol. 1 and Episode 456, Perona follows Zoro through the ruins of Kuraigana Island.
  84. One Piece Manga and Anime — Vol. 60 Chapter 592 (p. 2-3) and Episode 509, Perona learn of Moria's supposed death and is informed of its dubiousness by Mihawk.
  85. One Piece Manga and Anime — Vol. 60 Chapter 594 (p. 16) and Episode 512, Perona helps Zoro read the newspaper detailing Luffy's return to Marineford.
  86. One Piece Manga and Anime — Vol. 61 Chapter 597 (p. 5) and Episode 515, Perona tends to Zoro's wounds and is informed of the meaning of Luffy's message.
  87. One Piece Manga and Anime — Vol. 78 Chapter 779 (p. 3) and Episode 720, Perona laughs at Zoro being unable to drink alcohol until he masters Haki.
  88. Special Episode 3D2Y, Perona aids Luffy and Hancock in the fight against the World Pirates.
  89. One Piece Manga and Anime — Vol. 61 Chapter 602 (p. 4 & 13) and Episode 522, Perona informs Luffy, Zoro, and Sanji of the Marine ships gathering around the archipelago and helps them escape.
  90. One Piece Manga — Vol. 67 Chapter 658, cover story: From the Decks of the World Vol. 40.
  91. One Piece Movie — One Piece: Stampede.
  92. One Piece Manga — Vol. 107 Chapter 1080 (p. 9), Perona frees Koby.
  93. One Piece Manga — Vol. 48 Chapter 466 (p. 12), Perona is foaming at the mouth after being knocked out by Usopp.
  94. One Piece Anime — Episode 361, Perona is not foaming at the mouth after being knocked out by Usopp.
  95. One Piece Anime — Episode 456, Perona acts like a lonely child and tries to make Zoro her servant.
  96. One Piece Anime — Episode 456, Although Perona is in her ghost form, she raises dust upon falling on the ground.
  97. One Piece Anime — Episode 515, Perona scolds Zoro for continuing to train despite his injuries.
  98. One Piece Anime — Episode 522, Perona's hat bends against Zoro while she is scolding him.
  99. One Piece Manga — Vol. 61 Chapter 602 (p. 4), Perona's hat does not bend upon making contact with Zoro.
  100. One Piece Anime — Episode 524, Perona attacks Sanji for freaking her out and expresses sadness at Zoro leaving.
  101. One Piece Manga and Anime — Vol. 49 Chapter 475 (p. 17) and Episode 370, Nami wears a top with Perona's skull and crossbones.

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